?????????
(
?? ????
: Animalia) ??
?????? ????
?????????
????????
, ????????? ?
?????
?????????
Animalia
. ???? ?? ?????????? ??????
???? ??????
, ??????
????
, ?? ??? ????? ?????, ?????? (????) ?
????? ?????
, ???????? ???? ??? ????? ?? ???? ??????
blastula
, ?????? ?????? ???????? ???????. ?? ???? ???? 1.5 ??????
?????
?? ???? ????, ????? ??? ?????? ????? ??
?????
. ?? ???, ??? ??????
[
???? ????
]
?????? ??? 7 ?????? ???? ???? ????. ???? ???? ?? ???? ????? ???? ?? ?????, ??????? ??????. ???? ????? ??????? ????? ?? ????? 8.5 ???????? ????, ???????? ????? ?????? ????? 30?40
?????
?
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???? ?????
. ???? ????? ??????? ?????????? ?????? ?? ????? ????? ????, ??? ??????
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.
??? ???? ?????????? ??????? ???? ???????? ?
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(
Bilateria
), ??? ?????? ????
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, ??????
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?
???
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?
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. ???????? ????? ?? ????
Nephrozoa
, ??????? ??? ????? ??????:
?????? ???
(
Protostomia
) ?????? ?????? ???? ??
???? ??????
???
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(???? ?? ?
?????
),
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(????? ?? ??
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?
????????
),
???????
, ????; ??? ???
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(
Deuterostomia
) ????? ??
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; ????? ???????? ????? ?? ??? ??
Xenacoelomorpha
????? ???? ?? ???
Proarticulata
?????.
?? ?
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????
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?????? ???
.
?????????? ??? ???????? ???????, ???????? ???? ?????? ???? ???????:
- ?? ?????????? ??
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, ??????????
[1]
[2]
, ?????? ??????? ????? ???? ????? ????????? ?
?????????
. ?????? ??????, ??? ??????????, ??? ?????? ????????, ??? ????? ?
????
.
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) ?? ??????
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.
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, ??????? ????? ?????? ??????? ?????? ??????? ?? ??
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DNA
. ???? ??? ?????? ???????? ?????? ?? ??????
???????????
(???? ?????) ?????? ???????. ??? ??? ????? ???? ????? ???? ???? ?? ????, ???? ????? ????? ??????. ???? ?????? ?? ???? ????? ???? ?????? ???? ????? ????? ??????? ???????? ????? ???? ????? ?????. ??? ?? ?
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1973
?
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(????? ??????? ?? ????????) ?
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(????? ??????? ?? ????????? ?????? ????): ???? ???? ???????? ????? ?????? ????????, ????? ???????? ??????????? ?? ???????? (????? ????? ????)?
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???????? ????????
), ???? ?? ????, ?? ?? ?? ???? ???? ???? ????; ???? ??????? ?????? ??? ???? ????? ?????? ??????? ??? ?? ?????.
???????
, ????, ?????? ?????? ??????? ???????????, ?????? ?????? ????? ??????? ?
?????????
, ??? ????.
- ?????
???????
(Porifera) - ???? ???? ???????? ???????? ??????, ????? ??? ???????. ??? ?? ??????? ??? ???? ????? ????????? ???? ???? ??? ?????? ???????. ?? ????? ????? ?????? ????? 10,000 ????? ??????. ???????? ?????? ????? ???? ??????, ?? ?? ???? ??????, ??? ???? ?????? ??? ????? ?? ???? ??8,500 ???. ???????? ??????? ????? ????? ???? ??????? ????? ??????? ????? ????? ????????.
- ?????
????????
(Placozoa) ????? ?????? ???? ????? ????? ????? ???? ???? ????? ?????????, ????? ??????, ???? ????? ?????? ?????? (?????) ?? ??? ?????. ?????? ?? 3 ????? ??????: Trichoplax adhaerens ?2 ????? ?????? ??? 2017:
Hoilungia hongkongensis
,
Polyplacotoma mediterranea
[6]
.
- ?????
?????????
(Myxozoa) - ??? 1,300 ????? ?? ???? ???? ?????? ?????? ????? ????. ????? ??? ?? ????? ???? ?? ??? ?????? ????? ???? ???
????? ??????
??
???????
.
- ????????
???????
(Mesozoa)
- ?????
???????? ???????
(Eumetazoa)
- ?????
???????
- ???? ??? ??????? ?????
- ?????
??????
(Cnidaria). ????? ??10,000 ????? ?? ???? ???? ?????? ????? (????? ??? ????? ??? ????) ??????? ????? ???. ?????? ?? ?????? ??? ????
???????
(??? 6000 ?????),
?????? ??
,
?????? ????
,
?????? ?????????
?
???????
.
- ?????
????????
(Ctenophora). ????? ??150 ?????, ????? ???? ??????? ?????? ????? ?????? ???? ??????? ??????. ????????? ?? ???? ????? ??????? ????? ?????? ??????? ????? - ????? ?? ???? ????????? ??? 1.5 ?????. ??? ?????? ???? ????? ???? ????? ??????? ??????.
- ?????
???????
(Bilateria) - ???? ??????? ???????? ?? ???? ???? ??????? - ????? ?? ???? ???? ???? ??????? ????????, ?????, ?? ??? ??? (??????) ???? (?????), ??? ?? ?? (??? ????) ???? (??? ??????); ??? ?? ??? ?? ?? ????? ??? ????. ???????? ??? ?????? ??????? ?????? ???? ????, ?????? ?? ??? ??????? ???? ????????, ????????, ???????? ????????. ??? ????? ???????? ?? ???? ???? ??????????? (triploblastic), ????? ???? ????
????? ???
: ???????, ?????? ????????.
- ?????
???? ??? ??? ????
(Acoelomata)
- ?????
?????? ??????
(Platyhelminthes) ?????? ??????, ???? ??? ??. ????? ????? ??????: ????? ?????, ?????, ??????? ?????? Monogenea. ????? ?????? ???? ??????, ????? ?? ???? ??? ??????? ?????? ?? ??????. ??? ?????? ????????????, ????? ?????? ?????? ?????? ????? ????? 12 ?????.
- ?????
???? ??? ??? ????
(Coelomata)
- ?????
?????? ???
(Deuterostomia)
- ?????
????? ??
(Chaetognatha)
- ?????
????????
(Chordata) - ???? ???????? ?
??????
, ?
????
, ?
???????
, ?
??????
, ?
?????
??
??????
- ?????? ?
???
.
- ?????
????? ???
(Echinodermata). ???? ??6,500 ?????, ????? ???? ???? ????? ?????, ??????? ??? ???????. ???? ?????? ?????? ????? ???? ?? ????? ???? ?????? ?? ?????. ???? ????? ??? ???? ????? ????? ??? ??? ?????? ???. ?????? ?? ??? ??????:
?????? ??
,
????? ??
,
?????? ??
,
??????? ??
,
?????? ??
??????????? (????). ???? ????? ??? ???? ????? ????? ??? ??? ?????? ???. ??? ???? ??????? ???? ????? ???????? ???? ?????? ????? ???, ?? ?????? ????. ?????? ?? ??? ????? ????? ??? ????, ??? ??? ????? ?????, ????. ????????? ???????? ?? ??? ?????? ?? ??????? ????????????, ??????? ??????? ????????, ??? ?????? (coelom) ????????, ???? ????? ???????? ????? ??????? ???. ??? ?? ???? ?????? ??? ??????? ?????? ???????????. ?? ????? ?? ??? ??????? ???? ?????? ??????? ???? ?????? ?
?????????
.
- ?????
?????? ??
(Hemichordata)
- ?????
??????????
(Xenoturbellida) ????? ??? ????.
- ?????
?????? ???
(Protostomia)
- ?????
??????????
(Brachiopoda)
???? ???? ??????
????? ???? ??? ????? (??? ?????? ???????? ?? ????????) ???????? ??? ????? ?????, ????? ?????? ?????? ???? ??? ???? ??? ????? ??? ?????? ?????? ???????. ?????? ??????? ?????? ????? ???? ?????? ????? ?????? ????? ???? ?? ?????? ??? ????. ???? ???? ?????? ???? ???? ????? ???? ????? ?????? ????? ??????? ???? ?? ????????? ??????? ???, ??????? ?? ????????? ???? ????? ????? ?? ???? ?? ????????? ????? ???. ?????????? ???? ??? ????, ????? ???? ???? ???? ?? ??? ???. ?????? ?? ?????????? ??????? ?? ???????? ??? ????? ???? ????? ???? ?????? ??????? ????????, ?
????????
???????. ?????? ????? ??????, ?????? ??? ?????, ?????? ??????? ????? ??????. ?????, ?
???????????
, ??? ??????????? ??? ???? ????? ???? ?????? ?????? ????? ??????? ?????, ???????? ??????? ????????? ?????, ???? ????? ?????? ????? ?????.
- ?????
???????
(Bryozoa) ????? ??4,000 ????? ??????? ????, ???? ???? ????? ??????? ?
?????
?????? ??? ?????? ??????? ????? ????. ???????? ???? ????? ?????: ????? ??? ????? ?????? ???? ???????? ???. ?? ?????? ??????? ??? ??????? ?????? ?? ????, ??? ?????? ?????? ?????? ??? ????????. ?? ??? ?????? ???? ?????? ????? ?????? ???? ????????, ????? ??? ???? ??? ????, ????? ????? ???????. ??? ???? ???? ?? ?????? ??????, ?????? ????????, ??????? ?? ??? ????? ?????, ?????? ???? ?????? ??? ??????? ??? ???? ?????? ????? ??????. ??????? ???????? ???? ????? ???? ?????? ?????? ?????? ?? ???????, ??? ???? ?????? ?????, ????? ????? ?????? ?????. ???????? ??? ????? ???? ?? ????? ?????.
- ?????
????????
(Phoronida) - ????? ???? ?? ??20 ????? ??
???? ???? ??????
?????, ??????? ????????? (Phoronids),
????? ????
?????
???????
("???" ?? ?????? ???), ??????? ???????
?????
?????? ??????? ??????? ?? ?????? ?????. ?? ???? ?????? ?
??????????
??????, ????
????????? ??????
, ?????
????????? ?????????
, ????
???? ?????
??? ????? ?? 400 ?????. ????? ?? ????? ?????????? ??? ??2 ????????? ?????? ??1.5 ?????????, ?? ?? ????? ?? ?
???
????? ????? ??? 50 ?????????.
- ?????
??????????
(Entoprocta)
- ?????
??????
(Mollusca) ??? ??????? ???????? ???????? ????? ?? ???? ??????? ?? ??? ???? ????. ??????? ????? ?? ??100,000 ?????, ???? ????????, ??????? ???? ??????. ?????? ?? ??????? ????? ??? ?????? ??? ???? ?????; ???? ??? ??????? ?????? ??? ?????? ??????. ???? ?? ??????? ?? ??? ????? ??? ?????. ??? ?? ?? ???? ???? ?? ???
???????
, ?????? ???? ????? ??????????. ???????? ?????? ?? ??? ??????, ?????? ???? ???? ?????? ???? ?????. ??????? ?? ???? ?????? ???? ????? ?????? ?????? ?? ????: ?????? (
?????
), ?????? ????? ?????? ??????? ??????. ?????? ????? ????? ?????? ???? ???? ?????, ???? ?????? ????? ????? ???? ????? ???????. ??? ??, ?????? ??????? ?? ??? ?????? ???? ?????? ??????? ?????. ??????? ????????? ?????? ?? ?????, ?? ????? ?? ?? ??? ??? ????, ?? ?????? ???????.
- ?????
????? ?????
(Nemertea)
- ?????
??????????
(Priapulida)
- ?????
?????? ????????
(Sipuncula)
- ?????
????????
(Annechiuphora)
- ?????
?????? ???????
(Annelida) ??15,000 ????? ?? ?????? ???? ???? (??? ??? ????). ???? ?????, ???? ???? ???? ?????? ????? ???????. ?????? ??????? ?????? ??????? ?????? ??? ??? ??? ??? ??????? (????), ????? ????? ???????? (?????) ? ??? ?? ?????? ??????? ?? ????, ??? ???????? ????????? ??? ????? ??????. ?????? ????? ????? ?????? ???????:
?????????
,
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?
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.
- ?????
??????
(Echiura)
- ?????
???? ????
(Pogonophora)
- ?????
??????????
(Panarthropoda)
- ?????
????????????
(Arthropoda) - ???? ???? ???? ?????? ????
??? ??????
, ???? ????? ??????, ??????? ???? ??????? ??????. ???? ?????? ?????? ????? ????? ????, ???? ????? ????? ??1,134,000 ????? ????? - ???? ??? ???? ???? ????? ?????? ?????. ??80% ???? ???? ????? ?????? ???? ???? ?? ???????????? (?????? ???? ??????). ?????? ??????? ???????? ??? ?????? ?????? ???? ?? ???? ????, ????
?????
,
????????
,
???????
?
??????
. ????? ?????? ??? ?????? ?????? ????? ????? ????, ?? ??? ?????? ????? ???????. ????? ??????? ????? ??????, ????? ????? ?? ??????? ?????????? (~0.25
???????
) ??? ????? ??
???? ????? ????
?????? ???? ?????.
- ?????
????? ?????????
(Onychophora) ????? ??200 ????? ?? ???? ???? ??????? ???? ???? ????? ???? ???, ???? ????? ????????????? ???????? ???????. ?? ????? ???? ????? ?????? ??? ????? ????? ?? ???? ??? ??.
- ?????
?????? ???
(Tardigrada) ????? ??? 1,000 ????? ?? ?????? ?????? ??? 0.1 ??????? ??1.5 ??????. ?????? ??? ?????? ??? ?????, ??? ???? ???????? ??? ??????? ???, ?????? ??? ????? ???? ???????, ?????,
??????
???? ?
?? ??????
??? ??????. ?? ??????? ????? ??????? ???????? ????????? ??????? ???? ?????? (273- ????? ??????), ????????? ?????? ?? ??? 151 ????? ?????? ?????? ????? ???? ?? 1,000 ??? ?????? ?? ??? ????. ?? ??????? ????? ???? ??? ???? ??? ???, ??? ?????? ????? ???? ???? ????.
- ?????
???? ??? ??? ???? ?????
(Pseudocoelomata)
- ?????
?????????????
(Gnathostomulida)
??? ??? ????? ?????? ????? ?? ???? ?????, ?????
??????
?? ???? ????? ?????? ??? ????? ???? ????, ?????? ?? ?? ???? ????? ?? ??5 ?????? ?????:
????, ?????
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, ???? ??? ????? ????? ???????? ?? ?????? ?????
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????. 4 ??????? ???????? ??????? ???? ?? ???? ?
??????
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?
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, ?? ?????? ??????? - ????? ?????? - ??? ?? ?????? ????? ?????, ???? ????? ???? ???? ??????? ???? ?????? ????? ?????.
- ??? ????? ?
???
???? ?? ???? ???? ??? ?????? ???????????. ???? ???? ??? ????? ?????
????
, ??? ????? ???? ?
???
,
?????
?
?????
?????? ????? ????? ?????.
??????? ?????? ??????? ????? ????? ??????? ???,
???? ???? ?????
?????? ?
??????
, ?????? ?? ????? ???? ????? ????. ???? ?? ??? ???? ???? ???? ??
?????
, ??????? ?????? ???? ??
???? ??????
. ?
????? ???????
??? ????? ???? ??? ???? ????? ????? ???? ???? ?????. ?? ???, ???
????
???????? ????? ?????, ?????
???
, ???? ?? ???? ????? ???? ????.
??? ???? ???? ??20, ?? ?????? ?????? ?? ?????? ?????? ?"
??????
", ?????? ?????? ?? ??? ???? ????.
- ??? ????? ?
????? ??? ??? ?? ??? ???? ????
???? ???? ???? ?? ?????? ????? ?????? ????? ??????? ???????. ??????? ???????? ????? ??????? ?
?????? ????? ??????
??? ?????? ???? ??? ????? ??????? ??????? ????. ???? ?????? ??? ?????? ????? ??? ????? ?? ???? ???? ?????? ????? ?????? ??? ?????? ?? ?????
??????
.
??? ?????? ?? ?????? ???
?????
,
????????
??
???????
????? ???
????? ??
, ???? ????? ?????? ????? ?? ?? ??????? ???
??????
. ?? ???? ???? ?????? ????? ???? ???? ??????
[????? ?????]
???? ??? ?????? ???? ?????? ??????? ????? ???????? ??????.
- ??? ????? ?
???? ???? ????
????? ?
??????????
????????? ???????, ???? ???? ???? ???? ????, ????
???
,
?????
,
?????
,
?????
,
????????
????. ??? ???? ????? ????? ?????? ???? ?
????
?? ?????? ?
??????
. ???? ????? ??? ?????? ?????? ??? ????
????? ???
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?????
??
?????
.
??? ????
????? ???? ????? ?????? ?????? ?????, ???? ???? ????? ?
???? ????
?? ?? ????? ??????? ????? (????
??? ?????
?
??? ?????
). ?
??????
????? ????? ????.
???? ????? ????? ???? ????? ???? ?????? ??????, ????? ????? ?
??? ????
,
??????
???????? ?????. ????? ????? ???? ???? ??? ???????? ??? ????? ???? ?
???? ??????
?? ?????
???? ??21
.
- ??? ????? ?
????? ????? ???? ????
????? ????? ???? ???? (?? ?????????), ??? ???? ?
??????????
???????? ????? ???? ??? ???? ?? ??????? ??????????? ?? ???? ??????. ???? ?? ?????? ?????? ???????? ???? ????? ??????, ??????,
??????
???????????.
????? ???? ???? ???????
??????
?? ?????? ??
??????????
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??????????
???????? ??
????? ??????
[7]
.
-
???????? ????? ????
-
???? ?????
-
??? ???? ?????
-
????
-
??? ????
- ^
Avila, Vernon L. (1995). Biology: Investigating Life on Earth. Jones & Bartlett Learning. pp. 767?. ISBN
978-0-86720-942-6
.
- ^
"Palaeos:Metazoa". Palaeos. Retrieved 25 February 2018.
- ^
Saupe, S.G. "Concepts of Biology". Retrieved 30 September 2007.
- ^
Mentel, Marek; Martin, William (2010). "Anaerobic animals from an ancient, anoxic ecological niche". BMC Biology. 8: 32. doi:10.1186/1741-7007-8-32. PMC 2859860.
PMID
20370917
.
- ^
Bergman, Jennifer. "Heterotrophs". Archived from the original on 29 August 2007. Retrieved 30 September 2007.
- ^
Voigt, O; Collins AG; Pearse VB; Pearse JS; Hadrys H; Ender A (
23 ??????? 2004
). "Placozoa ? no longer a phylum of one".
Current Biology
.
14
(22): R944?5.
doi
:
10.1016/j.cub.2004.10.036
.
PMID
15556848
.
- ^
Braun, C., Stangler, T., Narveson, J., & Pettingell, S. (2009). Animal-assisted therapy as a pain relief intervention for children. Complementary therapies in clinical practice, 15(2), 105-109.