???????

??????? ??????????????? ?????????????????? ???????.

???????
?????????? ?????: Cryogenian ? Present , 670?0Ma
???????? ????????? e
???? :
?????????

??????

?????????? ( Animals ), ????????? ( Animalia ) ?????? ?? ?????? ????? ( Metazoa ), ?????? ?????????????? ???????????? ??????????????? ????????????? ?????? ??????????. ??? ?????????????? ?????????, ????????? ??????? ????????????? ?????? ?????????? ????????????????? , ???????? ?????????????????? , ???????????? ??????????????????? ( en:Motility ), ???????????????? ??????????????, ????? ???????????????? ??????????? ???????????? ???????????????????? ( en:Blastula ) ??????????????? ????????????? ???????????.

?????? ?????? 7 ??????????????? ??????? ??????? ???????????? ????????? ?????????????????????????, ??????????? 1.5 ??????????????? ??????? ???????????? ???????????????????. ???????? ?????? 1 ????????? ?????? ????????????. ???????????? ????? 8.5 ????????????? (0.00033 ????????) ????? 33.6 ??????? (110 ???) ??? ????????????. ??????? ???????? ???????????? ?????????????? ???????????????????? ???????? ???????????? ??????????. ???????? ???? ??? ( en:Food web ) ??????????? ??? ??????????. ????????? ???? ?????????? ????????????? ???????????? ??????? ?????????????? ???????????? ??????? ???? ???? ???????????? ?????????? ?????????? ??????? ???????????.

??????????? ???????? ????????????? ????????????????? ??????????? ??????. ???????? ???????????????? , ???????????? ??????? ???????????? ????? ?????????????? ????????? ?????????? ????????????????? ??????????????. ???????? ????????????????? ????? ????????????? , ??????? ??????????? , ???? ??? ????????????? ?????????? ????????????????????? ????????????.

????????????????? ??????????? ???????? ????????? ???????? ?????????? ?????? ??????? ?????????????????. ?????? 542 ????????? ???????????? ?????? ???????? ??????????? ?????????? ????, ?? ???? ????????? ????????? ?????????? ???????????? ???????????, ???? ???????????? ??????? ????????????. ??????? ????????????????? ??????? 6,331 ?????????? ???????? ??????????????; ??? 650 ????????? ???????????? ?????? ??????? ??? ??????? ?????????????????? ????????????????? ?? ??????????????.

????? ?????? [ ???? ]

"??????" ???? ?????? ???????? ?????? ?????? ???????? ???????????? ??????? ????????????. [3] ???????? ??????????? ????????? ?????????????? ??????? ????????????????? ????????????. [4] ??? ????? ??????? ??????? ?????? ?????? ????? ???? ?????? ????? ???????????? ??????? ??????? ?????????????. ?????? ?????????????? ????????????, ???? ???????? ??????? ?????????? ???????????? [ ???????? ???? ] ???????????. [5] [6] [7] [8] ??????? ????????? ( Kingdom Animalia ) ??????? ???? ???????????? ???????? ?????? ?????????? ??????????????? ????????????. "??????????" ???? ???? ?????? ??????? μετα (??????, "???????" ????? ?????????? ??????????????????) ??????? ζ?? (z?ia, ζ?ον z?ion "???????" ??????? ?????) ????? ??????????????? ????????. [9] [10]

???????? [ ???? ]

??? ??????????????????? ???????? ????????? ???????? ?? ???????? ?????????? ??????????. ?????????? ?? ?????????????? ??????????????????? ???????????. [11] ??? ???????? ?????? ??????????????? ??????? ???? ??? ?????? ????? ??????????????????? ?????????? ?????????????. ??? ???????? ???????? ???? ??????? ?????? ???? ????? ????? ???????????? [12] ??????? ?????? ???????? ???? ?????????? ??????????????? ????????? ????????? ???????????. [13] ?????? ????? ????????? ??????? ?????????? ?????????????????? ???????? ?????????? ??????????. [12] [14] ????? ???????????? ??????????? ???????? ??????????? ????? ?????? ????? ????????? ????? [15] ????? ?????????. ???? ????????????, ????? ???????????????? , ?????????? ??? ??????????? ??????? ???????????? ?????????????. ??? ?????????????? ?????????? ??????????? ?????????.

????????? [ ???? ]

?????????? ???????? ???????????? ????????????? ?????????????? ????????????????. ??????? ??????????? (???????????? ( Porifera ) ??????) ??????? ????????? ??????????? ????? ??????????????? ??? ???????????????? ?????. ????????? ????????? ?????? ???????????????????????? ?????? , ??????? ?????????? ?????????????? ?????????????????????????????? ?????? ??????? ???? ??????????? ????????. ??????? ??? ????????? ??????????? ?????????? ????? ?????? ?????? ????????????? ???????????????. [16] ???? ??? ??????????????? ????? ?????????? ????????????? (?? ?????? ???????????) ????? ?????????????????. ????????? ??????? ???????????? ??????????? ????????? ????????? ??????????????? ( eumetazoans ) ????? ?????????????????.

???????????????, ???????????? [ ???? ]

???????????? ???????  (1) ??????????? ????????? ??????????? ???? ??????? (2).

????????? ????? ???????????? ??? ??? ??????? ????????????????? ????????????. [17] ??? ??? ??? ????????????? ??????????? ????????????? ??????????. ???????? ????? ?????? ???????????? ?????? ????? ???? ???? ????????? ???????? ?????????????? ????????? ?????????? ?????????. ????????? ????????? ??????? ??????????? ??????? ???????????? ??????????? ??????????? ?????????, ??? ????? ??????????? ??????????????????.

???????? ????????????? ????????? ?? ?????????? ????????????????. ?????????????? ????? ??????????? ???????? ????????? ?????????? ???????? ????????????????. ??? ?????????? ??????? ????????????? ?????? ?????????. ?????? ??? ???????????????? ??????????? ( en:fragmentation ) ????, ?????? ?????????? ( en:Budding ) ???? ?????????? ??? ??????????????. [18] [19]

??? ?????????????? ?????????? ( en:Blastula ) ??????? ??? ????????? ????? ???????????. ??? ??? ???????????????, ?????? ?????????????????????????? ??????????. ?????????????, ????????????? ??? ????? ?????????? ???????? ?????? ??? ????? ???????? ????? ??????????. ?? ??? ???????????, ????????????? ??????? ???????? ??? ???????????????? ?????????. ??? ??????????? ???????? ??????? ??? ????????? ???????????????????? ???????????? ??????? ???????? ???????. ??????? ???????? ????????? ??????? ??????? ??? ???????? ???????? ???????? ???????????????.

???? ??????? ?????????? ?????? [ ???? ]

????????? ???? ???????????? ?????? ?????????? ?????????? ???????? ???????? ????????? ???????? ??? ???????? ???????????? ??????????????????. ?????????? , ???? ???????? , ?????????????? , ????????, ????????? ????? ????????????????? ????????????????????? ???? ?????? ?????? ????????? ????????? ???????????? ( en:Ditritivore ) [20] , ????????? ?????? ?????? ???????????? ????? ???????? ????????. [21] ????????????????????? ?????????? ?????? ????????????? ????? ?????????? ???????? ???? ????? ???????????. ??????? ??????? ????? ????????????? ??????????. [22]  

?????????? ?????? ???????????????? ???????? ?????? ??? ????????? ???? ?????????? ?????. [23] ????? ???????????? ??????? (????????????? ??? ?????????? ????? ???????), ??? ??? ?? (????????????? ?????????? ????????) ??????? ???????????. ?????????? ?????? ?????????????? ??????? ????? ??????????? ??????? ???????? ????????? ?????? ??????????? ???????. ????? ???????? ????????? ??? ????????? ????????. ????????? ??????? ??????? ?????? ????????? ?????? ??????????? ( detritivory ) ??? ?????. ?????? ????? ????? ?????? ?????? ????????. ??????? ?????? ???????????? ?????. ??? ?????????? ??????? ????????????? ??????? ?????????????? ???????? ??????. ???????? ???????????? ?????????? ??? ?????????? ?????????? ?????????. ???? ?????????? ???? ??????? ?????? ???? ??????????????????? ?????????? ??????? ???? ???? ??????? ????????? ?????????. ??????? ???? ??????????????? ??????????????? ???? ?????? ???????? ??????????.

??? ?????????? ???????? ???? ????????? ?????????? ?????????????. ????????? ???????, ??? ???????????????? ????????? ?????????? ?????????????? ?????? ???? ??????? ????????. ?????, ??? ?????????? ???????? ????? ????????????????? ????????? ?????????? ???????? ??????? ????? ?????????? ?????????????. [24] [25]

????? ??????????? ???? ????????? ????????????? ??????????? ???????????? ????????????? ????????? ??????? ????????????? ?????????? ??????? ???????????????. ??? ??????? ???????????-????????? ?????????????? ( en:Anti-predator adaptation ) ???????????????. [26] [27]

???? ?????????? ????? ??? ????????? ??????? ????????? ?????? ??????????. ????????? ????????????? ????? ??? ????????????? ??????????? ????? ????????????? ?????????? ??????? , ???? ???????????? ????????????. ??????? ???? (CO 2 ) ??????? ???? (H 2 O) ??????????????? ???????, ?????????????????? ????? ??? ??????? ???????? (C 6 H 12 O 6 ) ????????????? ??????????????? ????????? ???????? ???????? ????? ?????? (O 2 ) ?????????????. ???? ??????????? ???? ???????? ????????????? ?????????, ?????? ??? ??????????????. ?????????? ???? ????????? ??????????? (?????? ????????? ???????????? ????? ??? ?????????? ?????????), ?????????? ?????????????? ??????????? ??????????? ????????????????????. ??? ???????? ??????? ??? ??????????????????, ?????? ?????????????, ?????????????? ????? ??? ??????? ??????????, ??????????? ?????????? ???????? ??????? ???????????.

??? ????????? ??????? ???????? ????? [ ???? ]

?????????? ??????? ??? ??????????? ?????????????? ??????? ????????????????????? ????? ??????????????. ???????? ??? ????????? ?????? ????????????? ???????????? ??????????????????????? ??????, ??? ????????????? ?????????????? ???? ????????????? ????? ???????????????????? ?????????????????? ( flagellates ). ???????? ???????? ?????????? ??????????????? ??????? ??????????????? ??????????????????. ????? ???????????????????????, ????????? ??????? ??????? ????? ????????? ??? ??????? ??????????? ?????? ????????. ???? ???????????? ????????? ??????????? ???????? ?????? ?????? ????????? ?????????? (flagellum) ???????? ????? ??????????????? ??????? ???? ????? ????????. ??? ??????????????? ???????? ?????????? ??????????? ??????????.

?????????? ???????????????????????? ????? ????? ????????? ??????????? ??????????? ??????? ???????????? ????????????. ??? ?????? 610 ????????? ????????????? ??????? ????????? ??????? ?????????????. ???????, ??? ??????? ???????????? ????????????? ???????????? ?????. ??? ???? ??????? ????????????? ??????????????? ??????????? ??????????? ??? ??? ??????????? ??????????? ???????????? ??????????; ??? ?????????? ???? ?????????? ?????????????????? ??????. ??? ????, ???? ?????????? ??????? ????????? ?????? 542 ????????? ????????????? ???????? ??????????? ????????? ????????? ??? ????????????? ??????? ??????????. ??????????? ???????? ????? ????????????? ???? ??????????, ???????? ????????? ???????? ??? ????? ????????????? ??????????? ?????? ???????????? ?????????????? ?????????????? ??? ??????????? ??????????? ?????? ???????? ???????????????? ???????? ??????. ??????? ????????????? ????? ?????????? ???????? ????????? ????????? ???????? ??????????????????? ( paleontologists ) ??????? ??????? ????????????? ??????? ???????????? ??? ???? ??????????, ?????????? ?????? ????????? 1 ????????? ??????????????? ????????, ?????????? ???????????????? ???? ?????????????. ??????? ???????????? ????????? ??????? ??????? ????????? ????? ?????? ????????, ????????????? ????? ?????????????????? ????????? ????????? ???????????? ???????? ???????????? (?????? 5 ???? ?????) ??????????????? ???????????????? ?????? ???????? ????????????. [28] [29] [30]

?????????????? [ ???? ]

???? [ ???? ]

???????????? ????? ??????????? ( Balaenoptera musculus ) ?????? ?????????????? ?????????? ??????? ?????, ???? ??? 190 ?????? ??????? ????? 33.6 ??????? (110 ???) ??????? ??????. [31] [32] [33] ????????? ?????? ?????????? ??????? ?????????? ???? ( Loxodonta africana ). ???? ??? 12.25 ??????? [31] ????? 10.67 ??????? (35.0 ???) ???????. ?????? 73 ??? ???????? ???????? ??????????, ???????????????? ????? ???????????? ????????????. 39 ??????? ???????? ???????????? ?????? ?????????????? ?????????? ??????? ??????????????. [34] [35] ?? ?????????? ?????????. ?????? ?????????? ?????? ??? ??????. ???????? ???: ?????????? (?????????????? ?????????? ????????????) 20  ????????????????? ???? ??????. [36] ?????? ????? ????????? ????? ??????????? ?????. ??? ????????? ?????? ???? 8.5 ????????????????? ???? ?????. [37]

????????????? ????????? [ ???? ]

????????? ???????????, ??????? ??????????? ??????? ???????????? (???????????, ??????? [38] ??????? ???? [39] ) ??????? ???????? ???? (??????????? ???????? ?????? ????????? ???????? [40] ) ???????? ????????? ?????????????. [41] ?????????????????? ?????????????? ?????????????? ??????????? ???? ????????? ???????????????. ????? ????????????????? ?????????????? ???????????? ???????? ???????????? ?????????????????? ????????????? ???????????? ?????. ??????? ??????? ????????? ???????????? ??? ???????????????????. ?????? ??? ?????????? ?????????. ????????, ?????? 25,000?27,000 ?????? ????????????? ???????????????????. ????? ???????? ?????????????? ????? ????????????? ????????????? ?????????????? 10,000?20,000 ????????. [42] ???????????? ??????????? ???? ??????????? ???????????, ??????? ????????? ????? ?????????-?????? ???????????????? ?????-2011 ??? ?????? 7.77 ?????????? ???????????????. [43] [44] [a]

?????? ??????? ?????????????????? ???????????? ????? ???? ??????? ???????????? ?????? ????????? ????????
????????? wasp 1,257,000 [41] 1,000,000
(?????????) [46]
>40,000
(?????????????????) [47]
94,000 [38] ??? [39] >45,000 [b] [40]
????????? snail 85,000 [41]
107,000 [48]
35,000 [48] 60,000 [48] 5,000 [38]
12,000 [48]
??? [39] >5,600 [40]
?????????? green spotted frog facing right >70,000 [41] [49] 23,000 [50] 13,000 [50] 18,000 [38]
9,000 [50]
??? 40
(????????) [40] [51]
??????? ????????? 29,500 [41] ??? [52] ??? [39] 1,300 [38] ??? [39]

3,000?6,500 [53]

>40,000 [40]

4,000?25,000 [53]

????????????? 25,000 [41] ??? (????) [39] 4,000 [42] 2,000 [38] 11,000 [42] 14,000 [42]
????????????????? 17,000 [41] ??? (????) [39] ??? [39] 1,750 [38] ??? 400 [40]
???????????? Table coral 16,000 [41] ??? [39] ??? (???) [39] ??? [39] >1,350
(??????????) [40]
????????? 10,800 [41] ??? [39] 200?300 [38] ??? ??? [54]
??????????? 7,500 [41] 7,500 [41] ??? [39]
?????????? 6,000 [41] ??? [39] 60?80 [38] ???
?????????? 2,000 [41] >400 [55] 2,000 [38] ???
???????????? 1,335 [41] ??? [56]
(????? ?????????)
??? ??? ???
???????????????? 794 [41] ??? [56] ??? ???
?????????????? 430 [41] ??? [56] ???
??????????? 354 [41] ???
(????????? ???????) [56]
???
(??? ???????, ????????) [57]
??? ???
(?????????????) [56]
???
(???? ????? ????????) [56]
?????????????? 396 [41]
(30,000 ?????????????) [56]
??? [56] ???
???????????? 196 [41] ??? (??????) [56] ???
???????? 187 [41] ??? [56] ???
????????? 187 [41] ??? [56] ???
?????????? 186 [41] ??? [56] ???
????????????? 172 [41] ??? [56] ??? ???
?????????? 126 [41] ??? [56] ???
??????????? 107 [41] ???
???????????????? 97 [41] ??? (????) [56] ???
?????????? 30 [41] ???(????) [56] ???
??????????????? 29 [41] ???
???????????? 20 [41] ??? [56] ???
???????? 16 [41] ??? [56] ???
???????????????? 1 [56] ??? (????) [56] ???
??????????? 1 [41] ??? [56] ???
2013??? ??????? ??????????????? ?????????????? ????????????: 1,525,728

????????? ????????? [ ???? ]

???????????? ( Porifera ) [ ???? ]

?????? ?????? ???? ????????. ?????????????. ?????? ??????????????. ?????????????: ??? ???? ?????? ??????? ??? ???? ?????.

??????????? (????????????) ??????????? ??? ????????????? ??????? ???????? ???????????? ???? ???? ????? ??????????? ??????. ???? ??????????????? ???, ???????? ??? ?? ??????? ??????????? ?????????? ???????? ????????? ????????????????. ???????? ??????? ????????????. ????? ???? ???????????????? ???????? ?????????? ????????????????????? ??????????. ??????????? ???????????? . ??????? ???? ??????? ????? ?????????? ????? ???? ?????????????. ??????? 2008 ??? ??????? 21 ????????? [58] 150 ???????????? ??????????? ????? ????? ???? ??????????? ???? ????????????, ????????????? ???????? 21 ???????????, ??????????? ????????? ?????????? ????? ??????????????????.

???????????? ???????? ????????? ?????, ????? ??? ??????????? ?????????????????????. ??? ??????? ( ectoderm ) ??????? ????????? ( endoderm ) ???? ?????? ??????? ?????????? ?????????? ??????? ?????. ?????????? ??????? ??????? ??????? ??????? ??????????. ?????????, ???? ?????????? ??? ?????????? ???????? ( diploblastic ) ?????????? ????? ?????????????????. ??????????? ??????????????? ????????? ?????????. ????? ?????????? ?????????? ??????? ??? ????? ?????????.

?????? ?????????? ????????? ??????? ????????????? ?????? ???????????????. ???? ???????????, ??? ????????? ???????????????? ???????????. ???????????? ???? ??????????? ??????? ?????????? ?????????????? ????????????? ??????? ??????????. ?????? ????????? ???????? ??????????. ??????? ?????? ?????????? ???????????? ?????-??????????????????? ???????????. ????????? ???? ????????? ?????????? ???????????????. ??????? ??? ?????? ???????????? ???????????. ??? ???? ??????? ??? ???????????, ?????? ( coelom ) ?????? ?????????? ( pseudocoelom ) ??????? ??????? ?????? ???? ????????? ??????. ??????? ???? ?????????? ??????????????? ?????????????? ????? ? ???????? ????????? ??????????? ( echinoderm ) ????????? ??????????? ????????? ?????????. ??? ????????? ????????? ??????? ??????????????????? ???? ???????????? ??????????????.

??????????????? ?????? ??????? ????? ???? ????????? ????? ???????? ??????????????? ?????????? ????????????????. ????????? ????????????????? ??????? ??????????????? ??????? ?????? ??????? ???????????? ?????????????? ????????????.

????????????????? [ ???? ]

????????????????? ??????????????? ????? ????????????? ??? ???????????????? ??????? ?? ???????? ?????????????. ?????????? ??? ????????? ??????? ???? ?????. ???????, ????????????????? ????? ??????? (?????????????) ????? ?????????????????, ??????????? ?????? ??????????. ??????????????????? ??? ????????? ?????????. ????????????????? ??? ????????????????? ???, ???????????? ?????? ???? ??????????. ??????? ???????? ?????????????? ??? ???????? ??????????? ?????????????.

??????????? ??????????????????? ????????????????? ???????????, ????????????? ??????????? ???????? ????????????. ??????????????????? ??????? ?????? ??????????? ( Echinodermata ) ??????? ????????????????? (Chordate)??????. ????????? ????????? ??????????? ??????, ????????? ????, ???? ????????, ??????? ???? ?????????? ????? ?????????? ??????? ???????????. ????????? ???????????? ????? ??????????? ????????????????? ????????? ???????? ?????????????????????. ???????? ???? , ??????? ?????????, ????? , ??????? , ??????? ??????????? ?????? ????????.

??????? ?????? ????? ??????????? ?????????????????? ??????????. ????? ????????? ???????? ???????? ???????????? ?????????? ??????????.

??????????? [ ???? ]

??????-????? ?????????????

???????????????? ??????????????? ?????. ??? ?????????????? ?????? ?????????? ( ecdysis ) ????? ?????? ??????? ??????????? ???? ?????????????. ??????????? ??????? ????????? ???????????? ( Arthropoda ) ?????? ??????????. ????? ????????? , ?????????? , ???????? ??????? ???????? ?????????? ???????. ???? ??????? ????????????? ??????? ??? ????????????????? ???? ???????????? ???????????? ????????????? ??????????. ????????? ??????? ???????????? ???? ?????? ???? ??????????? ?????????????? ( Arthropoda ) ????????? ??????????. ??? ??? ?????????? ????????? ??????????.

???????????????? ??????? ?????? ???????????????????? ???????????????????. ??? ????????? ?????????? ??????? ????????????. ???????????????? ??????? ?????????????? ??????????, ????????? ???? ????????? ??????? ?????????????? ????????????????. ????????? ?????????? ????????????. ??????????? ?????? ?????????? ?????????, ??????? ??????????, ???????????, ??????? ??????????? ?????? ???????????? ?????????? ???? ????????? ?????. ???? ????????? ?????????? ( pseudocoelom ) ????? ?????????????? ??? ???????????? ???????? ??????????.

????????????????? ?????? ?????? ??????????? ??? ?????????? ?????? ????????? ????? ??? ??????? ??????????????????. ?????? ?????????? ?????????????? ?????? ???????? ???????????.

???????????? [ ???? ]

????????????? ???????????

???????????????? ???????????????? ( Platyhelminthes ), ???????????????? ???? ????????? ???????. ??? ??????????? ??? ??? ??????? ??????????? ???????? ???????? ???????????. ????? ??? ???? ??? ???????? ???????????? ??????? ??????????????????????? ??????? ??????? ??????????????. [59]

????????? ??????? ????????? ( flukes ) ??????? ??????????????? ????? ??????? ???????????? ???? ???????? ????????????. ??????? ????????? ?????????????? . [60]

??? ???????????? ????????? ???????????? ?????????? ??????? ???????? ?????? ( pseudocoelomate )???? ???????????. ???????? ????? ??????????? ????????? ??????????? ?????. ??? ????????? ????????????? ????? ?????????? ??????????. ???????? ????????????? ?????? ?????????-??? ?????????, ?????????????, ?????????????, ??????? ?????????? ?????? ??????????? ?????????? ????????. [61] ???? ????????? ??????? ???????? ??????? ??????????????? ????????? ??????????. ??????????? ??? ????????? ( Gnathifera ) ????? ?????????????????.

?????????????? [ ???? ]

????? ?????, ???????? ????????

?????????????? ???????????? ( Mollusca ) ??????? ????????????????? ( Annelida ) ???? ?????? ??? ??????????? ??????? ????????? ????????????????????. [62] [63] ??????? ?????????? ????????? ????????????? ???????????, ????????, ????????????, ??????? ?????????? ?????? ???????. ??????????? ???????????? ??????? ???????? ????? ??????????? ????????? ??????????????????. ???? ?????? ??????????? ???????? ??????? ????????? ?????????? ????????????? ??????? ???????????? ????????? ??????????? ???????????????. ????? ????????????????? ( Annelida ) ???????????????? ( Arthropoda ) ???????????????? ???????????????. [64] ????????? ???????? ???????? ???? ???????. [65]

?????????????? ????????? ?????? ??????? ?????????, ??????????, ??????? ???????? ????? ????????????? ?????? ?????? ?????? ??? ?????? ???? ??????? ?????????? ?????????????? ?? ????????? ????????????? ?????????????????. [66] ??? ?????????? ??????????????? ????? ?????? ?????????????? ?????. [67] ????? ??????? ??? ???????????? ???????, [68] ??? ??????????????? ???????????? ???????, ??? ???????????? ( Mollusca ) ??????? ????????????????????? ( Annelida ) ???????????? ??????? ??????????????. [69] [70] ???????? ?????????? ??????????? ?????????? ???????????? ?????? ??????? ???????, ?????????????, ????????, ??????? ?????????? ?????? ?????????? ?????? ???? ???????????? ???????? ????????????. [71]

?????? ??????????? [ ???? ]

???????????? ?????????? ?????? ????????????? ???????, ????????????? ??? ???? ?????????????? ?????? ?????? ????? ??????, ??????? ???????????? ??????????? ????? ?????????? ??????????, ??????? ?????????? ?????? ?????????????? ?????? ??????? ????????????? ??????????? ??????? ?????? ??????? ???? ??????????????? ????????? ??????? ??????? ???????? ??????????. ??????????? ????????????? ????? ????????, ???? ????????? ?????????? ???????????? ??????? ??????? ????????????? ????????? ?????? ???? ??????? ???????? ????? ??????????? ?????? ??????? ( metazoan ) ?????? ???????????? ???????????. ??? ???? ????? ??????? ???????? ????????????? ????? ???????? ?????????????? ???????????. ???????? ??????? ?????????? ???? ?????? ???? ???????. ????? ???? ??????? ????????? ???????????? ?? ??????????, ???????????? ??????? ????? ?????????? ??????????????. ???? ???????????????? ??????? ???????????? ????? ???????? ??????? ???? ????????? ????????. ???????????????????????? ???? ??? ???????????? ??????????? ??????? ???????? ?????????????? ??????????, ???????????????, ??????? ???????????? ??????? ???????????? ????? ????? ?????. ?????????? ???? ??????? ????????????????? ????????, ???????????????? ??????????? 1500 ?????????? ???????????? ??????? ???????????? ???????????, ???????????????, ??????? ??????????????????????? ??????? ?????????? ?????????????????????? ???????????????? ?????????????????. [72]

?????????? ???????? ???????? ???? ??????????? ??? ????????, ??????????? ??????? ????????? ???????????? ??????? ??????????? ????? ??????? ??????? ?????????????????? ??? ??????? ??????????? ???????? ????? ?????????????. [73]

?????????? ??????????????? ???????? ??? ?????? ????????????? ??? ( Mus musculus ) ??????? ??????????????? ( Danio rerio ) ?????? ???????.

???? ??????????????? ????? ?? ??????????? ?????? ???????

??????????? ?????? [ ???? ]

????????????????, ????????????? ???? ????? ??????????????? ???????????? ?????????? [?] ??????? ?????? (????????, ???????????????) ??????? ???????? ????? ??????????. ??????? ?????????? ??????????????? (???????, 2 ???????, ?????????? ??????????) ???????? ?????? ??????? (???????, 4 ???????, ????????? ???? ??????????) ??????? ???????????? (??????? ?????, ?? ???????, ????????? ?????) ??????????? (??????? ?????, ??????? ?????, ??????? ?????) ????? ???????????? ??????????? ??????????? ?? ????????????????. ????????????? ??????????? ?????????? ?????? ???????? ?????????? ??????????? ??????????????. ??? ????????? ??????, ??? ??????? ??????? ??????? ????????????? ?????? ??????????? ??????? ??? ???????? ???????? ?????????? ????????? ????? ?????????. ?????? ??? ??????????????? ????????????????? ?????????? ?????? ???? ???????????????. ??? ??????????? ??? ????????, ??????? ???????. [74]

1758-??, ?????? ????????? ???? ??????? ???????????? ????? ??????? ??????????? ?????????????. [75] ????? ???? ???????????, ?????????? ?????? ???????????? ?????????. ??? ?????????, ?????????, ???????, ??????? ??????, ??????? ??????? ???????? ?? ?? ????????????? ??????????????. ?????? ??????? ????????????? ?????????? ??????? ????????? ???? ??????????. ?????? ????????? ????????? ??????????????? ?????? ????????? ????????????? ??????. ??????? ????? ???????? ?????????? ?????? ????????? ???????? ????????? ??????????????????????. ?????? ???? ????????? ?????????? ???????????????????????? ???????????. ????????, ?????????? ??? ???? ?????????? ( protozoa ) , ??? ??????? ????????, ??????????? ?????????? ???? ???????????. ????? ??????? ??? ??? ???????? ???????????????.

????????? ????? ?????????????, ?????????? ?????? ?????????????? ??????, ???????? ( Vermes ), ????????? ( Insecta ), ??????? ( Pisces ), ???? ??? ???????? ( Amphibia ), ????????? ( Aves ), ??????? ????????? (Mammalia) ???? ????????????? ????????????????????. ??? ????? ????? ???????? ???????? ( Chordata ) ??????? ????? ???????????? ???????????????. ?? ??? ????????? ?????????????? ??????. ???????? ??????????? ???? ?????? ??????? ???? ???????? ????????? ???????????????. ??????? ??????????? ????? ??? ??????????? ??????.

??????? ?????????? [ ???? ]

  • ??????? ??????
  • ????? ????????
  • ???????????? ?????????? ????????
  • ??????????? ????????????? ??? ???????????? ????????
  • ??????

??????????? [ ???? ]

  1. The application of DNA barcoding to taxonomy further complicates this; a 2016 barcoding analysis estimated a total count of nearly 100,000 insect species for Canada alone, and extrapolated that the global insect fauna must be in excess of 10 million species, of which nearly 2 million are in a single fly family known as gall midges ( Cecidomyiidae ). [45]
  2. Not including parasitoid s. [40]

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??? ?????????? [ ???? ]