The Wayback Machine - https://web.archive.org/web/20090821021007/http://www.dinoweb.narod.ru:80/a_Theropoda.htm
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Theropoda
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adios85@mail.ru
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Theropoda (????????) - ???????? ????????? ?? ?????? ?? ??????? ????????, ? ??????????? ????? - ?????? (?? ??????????? ????????????????, ??????????, ???????????????? ? ??????????????, ???????, ????????, ???? ????????????????? ??? ?????????), ???????? ??????????????? ?? ?????? ?????. ????????? ????????????? ??????? ???? ??????? ??? ??????????? "????????????", ????? ?? ????? ??????? ? ???????. ????? ???????? ?????? ????????????? ????????????? ? ???????? ??????? - ?????????????? ????????? ? ??????????????? ????? ??????, ?????????? ??? ?????? ?????????? ? ?????? ??????? ?????, ????? ??????? ?????, ?????????? ?????? ?????? ????? ??????????? ? ?????, ? ????? ??????? ???????? (???????????) ? ?????????????? ?????????, ????? ?? ?????? ? ???????? ???.
???????? Theropoda ???? ??????? ???????? ?. ?. ?????? ? 1881?. ??? ?????? ????? ???? ????????? ? ???????? ? ???? ????????? Allosauridae, ?? ????? ???? ????????? ? ???? ?????? ? ????????? ?? ???? ????????? ????????, ??????? Megalosauridae, Compsognathidae, Ornithomimidae, Plateosauridae, Anchisauridae (??? ??? ????????? ?????? ????????? ? ?????????????) ? Hallopodidae (?????? ???????? ??? ???????????? ??????????). Theropoda ?????? ?????? ????????? ? 1866? ???? ?. ?. ????? ?????? Goniopoda ("?????????? ????"). ? ?????? 20-?? ???????? ????? ????????????, ??? ?. ???? ???????? ???????? Theropoda, ?????? ????? ????????? ?????? ?. ???? Saurischia, ??????? ?????? ?? ????????? Coelurosauria (???? ???? ???????? ??????????? ????? ?????? ???????) ? Pachypodosauria (???? ???????? ??????? ???????? ? ????????????), ??????? ?? ?????? ???????? Sauropoda, ??? ??? ???????????? ????? ??? ??? ????????? ??????? ????????? ??-?? ???????????? ?????????? ??????? ? ????? ???????????? ?????????? ? ?????? ???????????. ? 1922? ????? ? ????? ? ???????? ??????????? ??????? ????????? ??????????? ?? ?????????? ?????????? ? ?????????? ???????????? Goniopoda, ?? ??? ?? ???? ??????? ??????? ???????. ?????? ? 1956 ? Theropoda ???????????? ??? ??????, ?????????? ?????????? ??????????, ????? ?. ????? ??????????????????? ????? Saurischia ?? ??? ????????? - Theropoda ? Sauropoda. ??? ???????????? ? ????? ? ?????????????, ?? ??????????? Prosauropoda, ??????? ???? ???????? ? ???????? ??????????? ? Theropoda. ????? ????? ????????? ?????????? ??????? ?? Coelurosauria ? Carnosauria (???????? ???? ?. ???? ? ?????? 20 ????) ? ???????? ??? ???? ????????????. ?? ??? ?????????????? ???? ?????????? ??? ???????? ? ????????? ? 1969? ????????? ? ??????????, ??????? ?????? ???? ??????? ?? ? ????????????, ?? ? ???????????. ? ????? ? ???? ?. ????????? ? ????? 1970-? ?????? ????? ??? ???????????? ???????: Coelurosauria, Deinonychosauria, Oviraptorosauria, Carnosauria, Ornithomimosauria ? Deinocheirosauria. ? 1980-?? ?? ????????? ????????? ???? ????????????? ?????????? ??????? ?? ???????, ? ?????????? Tetanurae ? Ceratosauria, ? ???????? ???????? ?????? ????? ?????????? ? Maniraptora.
? ??????? ?????? ????????? ??? ?????????? ????????????, ??????????? ??????? ????????????, ? ????? ??????? ? ??? ???????????. ??? ?????????, ??????????? ???? ??????? ??????????, ??? ????? ????? ???? ??????? ? ????????? ???????????. ???????? ????????? ?? ?????????, ????????, ? ??????? ??????. ????? ??????????? ??? ??????? - ????????????, ?? ??????? (??? ???????? ????????? ????) ?????????: ??????????? ? ??????????? ???????? ? ??????? ???????? ??????, ???????????? ? ??????? ?????? ???, ? ????? ????? ??????????? ?????? ? ??????? ??????? ???. ??????????? ??????? ????????? ???????? ?????????, ?? ??? ?? ????? ????????? ???????? ???????????? ????????? ?????, ?????? ???? ? ????????, ??????????? ???? ?? ????? ???????? ?????? ???????? ???????? ?? ??? ??????? ?????.
?? 1980-? ?? ??? ???????? ???? ????????? ?? ??????? ?? ?????? ????????? ????, ????????? ????????????? (Coelurosauria - "???????????? ?????"), ? ??????? ????????, ????????? ???????????? (Carnosauria - "????? (?????????) ?????"), ??????????? ?? ?????? ????????????. ??? ?????????????? ???? ?????????? ? 1960-? ? 1970-? ?? ? ????????? ????????? ??????? ???????? ??????? ?????????. ??????? ??????? Carnosauria ?????? ????????? ????????, ?? ???????? ? ???? ????????????????? ??????? (??. ?????. ??????). ??????, ????? ????????????? ????????????? ??????? ???????????? ???????????? ??????? ?? ???????????? (Ceratosauria) ? ????? ???????? ?????? ??????? (Tetanurae).
?????? ?????? ??????? ????? ????? ????????, ????? ?? ?????? ??????? ?????????? ?????????? ???????. ? ???????????? ????? ?????????? ???????? ??????? ?? ?????? ???? 4 ?????? (?????????????, ?????????????, ??????????????????? ? ???????????), ?????????? ??????? ???????? ?? 10-14 ?????? ?????? ? 3-6 ???? ?????. ? ?????? ? ??????? ???? ????? ?? ???????? ????????? ????????? ????????????? ???????????, ??????????????????, ? ? ??????? ???? ?????? ???????? ???? ????????????.
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??? ????? ?????? ?????????? ? ???? ??????? ????? ??? ????? ??? ???.
?????????...
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???????? Theropoda:
????? ??????????????????; ??????? ????????? ???????????? ?????????????? ? ???????? ???????, ???????? ??????????????? ?????; ??????? ??????? ??????? ?? ???????? ? ???????? ????????????? ? ?????????????? ?????; ??????????????? ????????? ?????? ???????????? ?? ????????????????? ??????? ?????????????? ????; ??????? ????? ???????????? ?? ??????? ??????; ????? ???????????? ????? ? ??????; ??????? ????? ?????? ?? ?????????? ?? ?????????? ?????; ?????? ??????????? ?????? ? ??? ? ????-?????????? ????? (?????????????? ??????????? ????????? ? ??????????? ?????; ???? ????????? ? ?????????? ????? ? ????????? ????? ??????; opisthotic ? ??????? ?????????? ????? ?????? ?????????; ???????????? ?????????? ?????????????? ??????????; ???? ??????? ?????????????? ?????????; ?????? ????? ??????? ????? ????????? ????? ?????????? ??????; ??????? ????-???????????? ???; ????????? ???? ????? ??????????? ?? ????? ?? ???????? ??? ? ???????? ???????; ??????? ????? ? ???????? ??????; ???? ?????? ????? ?? ???????, ?????????, ? 2 ???????????? ???????; ?????? 23 ?????????????? ????????; ???????? ??????? ????? ????? ????????? ?????? ????????? (???????? ? ????????? ??????????? ?????); ??????? ????????????? ????????? ?????????; ????? ??????????? ?????? ????????????????, ????? ??? ????? ? ????????; ??????????? ????? ???? ??????? ? ???????; ???????? ????? ???????; ???????? ????? ?????????? ???????? ??? ?????? ????? ?????; ??????????? I-III ?????? ??????????; ??????? ???? ??????????; ?????? ??? ?????????? ?????????? ? ?????? ??????? ?????; ?????, ???????? ?? ?????, ???????, ???????????? ? ??????; ??????????? ????? ??????? ? ??????? ????????; ??????????? ?????????? ????? ???????? ????? ("???????? ???????"); ???????? ????? ????? ????????? ? ??????????, ???????? ????? - ??????????? ??? ??????; ???????? ????? ? ????? ?????????? ?????????; ??????????? ????? ????????? ? ?????????? ???????? ???????????? ????? ?????; ?????? ????? ????? ????? ????????? ? ???????? ???????? ???????????? ?????; ??????????? II ? IV ????????????, ? ??????????? III ??????????? (??????????????????? ????????? ??? ?????????? ?????? ?? ????? ????????????).
????????????? ??????????? ???????
????????
Dinosauria
___?????
Saurischia
______????????
Theropoda
_________??????????
Herrerasauria
____________?????????
Guaibasauridae
____________?????????
Herrerasauridae
_________??????????
Coelophysia
_________??????????
Ceratosauria
_________??????????
Tetanurae
Theropoda ??????????????
??????????? ? ??????? ???? ??????? ?? ??????? ????? ???????? ?? ????? ??????? ?????????, ????? ??????? ????????. ?????? ?? ??? ???????? ???? ?? ?????, ? ???? ????????? ?????????, ????????? ?? ?????? ???? ??????? ????????? (????
Troodon
) ? ??????? "??????????", ??? ?????? ??????????, ??????? ???????????? ????????????????? ???????? ???????? ?? ?????? ???????? ?????????.
Agnosphitys cromhallensis
(???????? ???????????)
Fraser, Padian, Walkden, and Davis, 2002
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?????-??? (??????? ?????)
??????
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??????? ?? ????? ????? ???????? ????? ????, ??????????????? ?????, ???????? ? ???????? ?????. ??? ?????? ???????? ????? ???? ???????, ??? ????? ???????? ??????????? ???????????? ?????????, ? ????? - ??????? ?????????. ????????? ????????????? ??????? ??? ? ????? ?????? ????????, ?????? ??????? ??? ???? ????? ???????????, ??? ???????? ? ???????????????.
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Lametasaurus indicus
(N.D.)
(?????????? ???????)
Matley, 1921
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????????? (??????? ???)
?????
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???? ??????, ?? ?????????, ??? ????????????? ??????????, ?? ??????? ??? ??????????? ???????????? ?????????, ???????? ? ?????????????? ????????? - ??????????? ??? ??????????????? ?????????. ??? ?????, ?????? ????????, ???????????? ?? ???? ????? ????? ?????????? ? ???????????? ??? ????????????. ?????????????? ???????? ???????? ???????, ??????????? ?????, ??????? ???????? ?????, ??????? ?????? ???????? ??????????? ???????? ?? ????????? ???????????
Rajasaurus
. ???????? ???? ?? ???????? ?????? ? ?????, ??????? ???????? ??????????????? ????????? ?????????? ? ?????? ????? ???????? ???????.
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Thecospondylus horneri
(N.D.)
(??????????? ???????)
Seeley, 1882
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?????? (?????? ???)
??????
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???????? ???????? "???????? ? ??????". ??? ???????????? ??????? ????????, ?????? ?????, ???????, ????????? ?? ??????????? ?????? ?????????? ?????????, ??????? ??????????? ?????? ???????? ?????????. ??????? ???? ????? ????,
T.daviesi
????? ??? ??? ??????????? ???
Thecocoelurus
.
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Herrerasauria
????????????? ????????? ? ??? ????? ??????????? ????? ???????, ? ??? ?????? ? ?????? ???? ????????, ? ?? ? ?????? ??????????. ? ??? ?????? ????? ???? ?? ??????? ????????? ?????? ???????, ?? ????????????? ???????? ???????? ?????? ???????? ?? ????????? ????? ???????. ??? ???????? ????????? ? ???? ??? ??????????? ???????? ????? ??????? ???????, ??? ? ??????????? ???? 4-????? ????? ? 5-? ?????????????, ? ????? ?????? ????? ??????? ??????? (?? ???????? ??????????? ????????? ? ??????? ?????? ???? ??????? ???????, ? ?????????? ???????). ?????? ????????? ???????- ?????????? ????? ???????? ?????, ????????? ??? ? ?????????, ? ????? ??????? ?? ????? ?????????????? ? ???????????????? + ????. ?? ?????????? ??????? ?????? ???????? ??????? ? ???? ?????? 2 ?????????? ???????? - ????? ??????? ????? ???? ??????????.
Herrerasauria - ????? ?????? ?????????, ?????? ??????????????? ??????? ????? ??????? 228 ???. ??? ?????, ?? ??? ???????? ? ????? ?????????? ???????. ??? ???? ?????? ? ??????? ???????? ???????, ????? ????????? ? ?????? ?? ?????????? ???????? ????????????? ??????? ? ????? ???????.
???????? ?????????????: ???????? ???????? 2-5 ?????? ??????; ???? ???????; ???? ???????????????????? ??????????; ??????????? ???????: 15+9+2+ ~50 (???????? ???????? ?? ?????? 2 ??????????? ?????????? ????????); ??????? ???????? ? ??????? ????????? ??????????; ???? ?????????????? ?????????? ????? ???????? ??????????; ?????? ?????? ??????????????????; ????????????? ?????????? ????????? ????????? ????????????? ?? ????????? ????????; ??????? ????? ? ?????????????; ?????? ???????? ???????????? ???????? ??????? (??? ???????????? ?????); ?????????? ?????; ????????? ? ????? ?????? ????? ?????? ?????????, ? ?????? ?????? ? ?????? ?????? ?????????? ? ? ???????? ???????; ?????????? ??????? ???? ??????? ?????????????; ???? brevis ??????????????; ??????? ?????????? ?????????? ???????? ?????; ????? ???????????, ? 5 ????????; ??????, ??????, ?????? ? ????????? ?????? ????? ? ???????????? ???????.
Guaibasauridae
????????????? - ????????? ??????????? ???????? ?? ???????? ?????? ???????? ? ?????????, ??? ?????? ???????????? ??? ?????? ???????? ???????. ????????????? ??? ????????? ???? ??????? ??? ?????? ???? ? ????, ?? ????? ????? ??????? ????????? ????? ? ??????? ????????????? ?????????, ???????? ? ???? ???????? ???? ??????? ????? ???????? - ??????? ? ?????????????. ????????? ????????????? ?????????? ????? ??? ???????? ?????????? ???????? ?? ??? ?????????, ???? ??? ???????, ??? ? ????????. ??? ?? ?????, ? ?????? ??????? ???????? ?????? ?????????????? ? ??? ? ???. ???? ?? ????????? ??????, ????????????? ??? ???? ???????? ?????? ????????????? ???????, ?????? ?????????????.
Herrerasauridae
Staurikosaurus pricei
(???????????? ??????)
Colbert, 1970
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?????? ?????? (??????? ?????)
????????
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???????????? - ???????????? ? ?????? ???????????, ????????? ?? ???????? ?????????? ??????????? ??????????, ? ???????? ? ????????? ??????? ????????????? ????? ?????????? ? ?????????????, ? ?? ???????, ? ?? ????????. ?????? ????????? ????????????? ??? ????????? ? ???????? ????? ????? ? ??? ?????? ?? ??? ? ????? ????????. ??? ???????? ????????? ? ????????? "????? ?????", ??????? ?????? ? ????? ?????????. ???????? ???????? ????????????? ????? ???????? ? ???, ??? ???? ???? ??? ????? ???????? ??? ?????? ??????? ? ??? ??????, ???? ?? ?????? ? ?????, ??? ???????? ?? ???????????. ??? ???? ?? ????? ?????? ??????????, ???????? ?????????? ??????? (???????? 2 ????? ?????? ? ????? 30 ?? ?????). ????? ????????? ???????? - ??????? ??????????? ???????, ?????????? ? ?????? ???????. ?? ????? ???? ?????? 2 ?????????? ????????, ??????? ???? ??? ????, ? ??????? ? ?????? ????? ??? ????????????.
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Herrerasaurus ischigualastensis
(??????????? ???????????????)
Reig, 1963
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?????? ?????? (??????? ?????)
?????????
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???? ???? ?? ????? ?????? ??????? ???????? ?? ?????? ???????? ?????????? ????????. "???????????" ?? ??? ?????? ??????? -
Ischisaurus cattoi ? Frenquellisaurus ischigualastensis
, ?????? ????? ???????? ?? ?????? ????????? ????????? ? ?????????????? ????????? ?? ? ?????????????, ? ??????? ??? ???? ???????? ????????. ?????????? ??????????? ?? ?????? ?????? ???????? ?? ????? ??????? ???????, ?????????? ????????? ????????, ? ?????????? ????????? ? ????? ?????? ???????.
???????? "???? ???????" ?????????? ?? ????????? ?????, ??? ?????????? ?????? ?????????? ??????? ????? ?????. ??? ????????? ????? ??????? ???? ?????????? ?? ?????? ?????? ????????? ? ?????????? ????????? ???????? 228 ????????? ???. ??? ? ??? ???? ??????? ?????? ? 1963?. ?? ?????????? ?????? ??? ????????????? ???????????? ?????????? ??????? ? ????? ? ???, ??? ???????? ???? ????????????? ???????? ?? ????? ????????????? ???????? ? ????? ???? ??????????? ?????????, ??????????? ???????????????, ??????????? ????????? ??? ?????? ?? ??????????. ??????, ? ????????? ? 1988? ??????? ? ?????? ????? ??? ? ???????? ???????????????? ??? ??????? ???????????? ????????. ???? ??? ??????? ????????, ? ?????? ? 3-4 ????? ? ????? 200-250 ??. ??????????? ????? ????? ? ??????? ?????, ???????????? ?????????? ????? ?? ???? ??????????????, ??????? ????????????? ????? ??????? ???????. ???? ???? ???? ??? ????????? ? ??????, ??? ?? ??????? - ??????? ? ???????. ????? ??????? ? ???????? ???? ??? ?????????????? ????????? ? ???? ?????????, 1 ?? ??????, ????? ??????????????? ?????????, ??? ?????? ??? ????? ??????????? ????????? ??? ??????. ?????? ?????????? ? ?????? ??????? ????????? ??????? ?? ???? ? ??????, ????????? ????. ????? ????????????? ???????? ??? ??????????, ?? ????????? ?????????? ? ????????? ??????. ??????? ???? ????????? ???????? ???????????? ?????? ??? ?????????? ????. ??? ?? ???? ???????? ? ??????. ???????? ???? ???????????? ???????????? ???????? - ?????? ???????? ????? ??????, ?????? ????? ???? ???????. ?? ?????? ??? ?????? ? ??????? ????? ????? ????????? ? ?????? ????? ??? ??????? ??????, ? ????????? ? ????? ?????? ???????????? ?? ???? ???? ???????? ??????? ??? ??????. ??????? ?????? ???? ? ????????? ??????? ? ???????? ??????? ???????? ?????? ??????? ??????. ???????? ??????????? ?????, ????????? ??? ????????????, ????? ???????? ???????? ?? ????????????????? ? ????.
???? ????? ?????????????? ?????? ?????. ??? ??? ??????? ??????????? ??????????, ?? ????? ???????? ?????????? ???????, ??????? ???? ???????? ???????. ??????????? ????? ?????????????? ?????? ????? ??????????? ??????????, ? ???????? ????? ?????????? ?????, ??????? ????, ??? ???????????? ? ???????????? ? ????. ????? ????, ????? ???????? ????? ????? ????? ???????, ??? ??????? ? ????? ??????????? ???????, ? ?????? ????????? ????? ????????-???????? ????? ? ???? ???????? ?????.
???? ?????? ???????????? ?????? ???????? ???????????, ?? ??? ???????? ????? ?????? ???????????? ??????????? ???, ? ??? ????????????????? ?? ???????. ? 1970 ? ???? ??????????????? ??? ??? ????????????, ? 1972 ? ?. ??????? ???????? ??? ????????????????? ??? Saurischia. ????????? ???????????? ???????? ???????????? ? ????????????? ?????????? ????? ?????????? ? ???????????, ? ????? ?????? ??? ?????? Dinosauria. ? ??????????? ??????? ?????? ????? ??????????? ??? ????????????? - ?????????????? ? ??????????? ????????? ? ?????? ? ?????? ????????, ????? ???????????? ?? ?? ??????? ? ?????????????. ???? ??? ????????, ?? ????????, ?????????????? ? ????????? ????????? ??? ??????, ??? ??????????????, ????? ???????, ? ??? ??? ????? ?????????? ??????? ????????? ???????????? ????????????, ???? ????? ???????????? ?????????? ??????? ? ???????? ?????????? ???????. ??? ????????????? ?????????????? ?????????????? ?????????? ??????, ??????? ????? ???? ????????? ?????? ????????, ?? ??????? ???????? los Rastros ? ?????????, ??????? ???????????? ???????????? ?? 3-5 ????????? ???.
???? ?? ?????, ??????????? ??? ?????????? ????????, ??????????? ?? ?????? ? ??????? ????????. ??, ????????, ??????? ??????? ??????????? ???? ???????????, ?????? ? ?? ??????? ??????? ????? ???????? ??????? ???? ?????????? ? ???????????????? ?????????. ?? ?????? ????????????, ????????, ????????? ?????????? ?????????? ???? ?????????, ??? ??? ?? ????? ?? ??????? ???? ??????? ????? ?? ?????.
??????: ??. ? ????? ????????
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??????? ?????? ? ??? ?? ??????, ??? ? ???????????, ? ??? ???? ??????? ? ????? ??????. ????? ???????? ??? ??????? ???????? ?????????? ????????????.
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????????? ???????? ????? ????? ????? ???????? ????? 56 ??, ? ??? ???????????? ?????????, ??????? ???? ??????? ?????? ???????????? - ?? 5-6 ?????? ?????? ? 350 ?? ?????.
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??????:
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