Hadrianus (turtle)

Hadrianus ; Temporal range: 55.4?33.9 Ma Pre? ? O S D C P T J K Pg N Eocene
	Hadrianus majusculus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade :	Pantestudinoidea
Superfamily:	Testudinoidea
(unranked):	† Pantestudinidae
Genus:	† Hadrianus ; Cope , 1872
Species
	H. corsoni ; H. majusculus ;

Hadrianus is an extinct genus of tortoise belonging to the Testudinidae ^[1] found in the United States , the Yolomecatl Formation of Mexico , the Alai Beds of Kyrgyzstan and Spain ^{[
citation needed
]} and believed to be the oldest true tortoise known. ^[3] The genus is thought to be closely related to the genus Manouria . ^[3] The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe. ^[3] Evangelos Vlachos (2018) reassessed the North American species attributed to the genus, and determined only two as accepted namely H. corsoni & H. majusculus . The remaining species were identified as either junior synonyms, moved to other genera or considered nomen dubium do to incomplete fossils. ^[4]

Taxonomy [ edit ]

Hadrianus corsoni (Leidy, 1871)
- Synonyms Emys carteri Leidy, 1871 ; Testudo hadriana Cope, 1871 ; Hadrianus quadratus Cope, 1871 ; Hadrianus octonaria Cope, 1871 ; Hadrianus tumidus Hay 1908 ; Hadrianus robustus Gilmore, 1915 ; Hadrianus utahensis Gilmore, 1915 ; Geochelone gilmorei Auffenberg, 1974
Hadrianus majusculus Hay, 1904

Nomen dubium [ edit ]

"Hadrianus allabiatus" Cope, 1871

Moved species [ edit ]

" Hadrianus schucherti " Hay, 1899 moved to Cymatholcus schucherti (Hay, 1899)

References [ edit ]

^ ^a ^b ^c "Fossilworks: Hadrianus" . fossilworks.org . Retrieved 17 December 2021 .
^ E. D. Cope. 1872. Second account of new Vertebrata from the Bridger Eocene. Proceedings of the American Philosophical Society (separate) 1-3
^ ^a ^b ^c Ehret, Dana Joseph 2004 "Skeletochronology as a method of aging Oligocene Gopherus laticuneus and Stylemys nebrascensis , using Gopherus polyphemus as a modern analog" Thesis, University of Florida.
^ Vlachos, E (2018). "A Review of the Fossil Record of North American Turtles of the Clade Pan-Testudinoidea" . Bulletin of the Peabody Museum of Natural History . 59 (1): 3. doi : 10.3374/014.059.0101 . ISSN 0079-032X . S2CID 214641639 .

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[PBDB-1] "Fossilworks: Hadrianus" . fossilworks.org . Retrieved 17 December 2021 .

[2] E. D. Cope. 1872. Second account of new Vertebrata from the Bridger Eocene. Proceedings of the American Philosophical Society (separate) 1-3

[Dating-3] Ehret, Dana Joseph 2004 "Skeletochronology as a method of aging Oligocene Gopherus laticuneus and Stylemys nebrascensis , using Gopherus polyphemus as a modern analog" Thesis, University of Florida.

[Vlachos2018-4] Vlachos, E (2018). "A Review of the Fossil Record of North American Turtles of the Clade Pan-Testudinoidea" . Bulletin of the Peabody Museum of Natural History . 59 (1): 3. doi : 10.3374/014.059.0101 . ISSN 0079-032X . S2CID 214641639 .

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