Plant or grouping of plants selected for desirable characteristics
A
cultivar
is a kind of
cultivated
plant that people have selected for desired
traits
and which retains those traits when
propagated
. Methods used to propagate cultivars include division, root and stem cuttings, offsets,
grafting
,
tissue culture
, or carefully controlled seed production. Most cultivars arise from deliberate human
manipulation
, but some originate from wild plants that have distinctive characteristics. Cultivar names are chosen according to rules of the
International Code of Nomenclature for Cultivated Plants
(ICNCP), and not all cultivated plants qualify as cultivars. Horticulturists generally believe the word
cultivar
[nb 1]
was coined as a term meaning "
cultivated variety
".
Popular
ornamental plants
like
roses
,
camellias
,
daffodils
,
rhododendrons
, and
azaleas
are commonly cultivars produced by
breeding and selection
or as
sports
, for floral colour or size, plant form, or other desirable characteristics.
[1]
Similarly, the world's agricultural
food crops
are almost exclusively cultivars that have been selected for characters such as improved yield, flavour, and resistance to disease, and very few wild plants are now
[
when?
]
used as food sources. Trees used in
forestry
are also special selections grown for their enhanced quality and yield of
timber
.
Cultivars form a major part of
Liberty Hyde Bailey
's broader group, the
cultigen
,
[2]
which is defined as a plant whose origin or selection is primarily due to intentional human activity.
[3]
A cultivar is not the same as a
botanical variety
,
[4]
which is a taxonomic rank below
subspecies
, and there are differences in the rules for creating and using the names of botanical varieties and cultivars. In recent times,
[
when?
]
the naming of cultivars has been complicated by the use of statutory
patents
for plants and recognition of
plant breeders' rights
.
[5]
The
International Union for the Protection of New Varieties of Plants
(UPOV ?
French
:
Union internationale pour la protection des obtentions vegetales
) offers legal protection of plant cultivars to persons or organisations that introduce new cultivars to commerce. UPOV requires that a cultivar be "distinct", "uniform", and "stable". To be "distinct", it must have characters that easily distinguish it from any other known cultivar. To be "uniform" and "stable", the cultivar must retain these characters in repeated propagation.
The naming of cultivars is an important aspect of
cultivated plant taxonomy
, and the correct naming of a cultivar is prescribed by the Rules and Recommendations of the
International Code of Nomenclature for Cultivated Plants
(ICNCP, commonly denominated the
Cultivated Plant Code
). A cultivar is given a cultivar name, which consists of the scientific
Latin
botanical name followed by a cultivar
epithet
. The cultivar epithet is usually in a
vernacular
language.
Etymology
[
edit
]
The word
cultivar
originated from the need to distinguish between wild plants and those with characteristics that arose in cultivation, presently denominated
cultigens
. This distinction dates to the Greek philosopher
Theophrastus
(370?285 BC), the "Father of Botany", who was keenly aware of this difference. Botanical historian Alan Morton noted that Theophrastus in his
Historia Plantarum
(
Enquiry into Plants
) "had an inkling of the limits of culturally induced (
phenotypic
) changes and of the importance of genetic constitution" (
Historia Plantarum
, Book 3, 2, 2 and
Causa Plantarum
, Book 1, 9, 3).
[6]
The
International Code of Nomenclature for algae, fungi, and plants
uses as its starting point for modern
botanical nomenclature
the
Latin
names in
Linnaeus
' (1707?1778)
Species Plantarum
(tenth edition) and
Genera Plantarum
(fifth edition). In
Species Plantarum
, Linnaeus enumerated all plants known to him, either directly or from his extensive reading. He recognised the rank of
varietas
(botanical "variety", a rank below that of
species
and
subspecies
) and he indicated these varieties with letters of the
Greek alphabet
, such as α, β, and λ, before the varietal name, rather than using the abbreviation "var." as is the present convention. Most of the varieties that Linnaeus enumerated were of "garden" origin rather than being wild plants.
[7]
In time the need to distinguish between wild plants and those with variations that had been cultivated increased. In the nineteenth century many "garden-derived" plants were given horticultural names, sometimes in Latin and sometimes in a vernacular language. From
circa
the 1900s, cultivated plants in
Europe
were recognised in the Scandinavian, Germanic, and Slavic literature as
stamm
or
sorte
,
[8]
but these words could not be used internationally because, by international agreement, any new denominations had to be in Latin.
[9]
In the twentieth century an improved international
nomenclature
was proposed for cultivated plants.
[10]
Liberty Hyde Bailey
of
Cornell University
in
New York
,
United States
created the word
cultivar
in 1923 when he wrote that:
The cultigen is a species, or its equivalent, that has appeared under domestication ? the plant is cultigenous. I now propose another name, cultivar, for a botanical variety, or for a race subordinate to species, that has originated under cultivation; it is not necessarily, however, referable to a recognized botanical species. It is essentially the equivalent of the botanical variety except in respect to its origin.
[2]
In that essay, Bailey used only the rank of species for the cultigen, but it was obvious to him that many domesticated plants were more like botanical varieties than species, and that realization appears to have motivated the suggestion of the new category of
cultivar
.
Bailey created the word
cultivar
. It is generally assumed to be a blend of
culti
vated
and
var
iety
but Bailey never explicitly stated the
etymology
and it has been suggested that the word is actually a blend of
culti
gen
and
var
iety
.
[11]
The
neologism
cultivar
was promoted as "euphonious" and "free from ambiguity".
[8]
[nb 2]
The first
Cultivated Plant Code
of 1953 subsequently commended its use, and by 1960 it had achieved common international acceptance.
[12]
Cultigens
[
edit
]
The words
cultigen
and
cultivar
may be confused with each other. A
cultigen
is any plant that is deliberately selected for or altered in cultivation, as opposed to an
indigen
; the
Cultivated Plant Code
states that cultigens are "maintained as recognisable entities solely by continued propagation".
[13]
Cultigens can have names at any of many taxonomic ranks, including those of
grex
,
species
,
cultivar group
,
variety
,
form
, and cultivar; and they may be plants that have been altered in cultivation, including by
genetic modification
, but have not been formally denominated.
[14]
A cultigen or a component of a cultigen can be accepted as a cultivar if it is recognisable and has stable characters. Therefore, all cultivars are cultigens, because they are cultivated, but not all cultigens are cultivars, because some cultigens have not been formally distinguished and named as cultivars.
Formal definition
[
edit
]
The
Cultivated Plant Code
notes that the word cultivar is used in two different senses: first, as a "classification category" the cultivar is defined in Article 2 of the
International Code of Nomenclature for Cultivated Plants
(2009, 8th edition) as follows:
The basic category of cultivated plants whose nomenclature is governed by this Code is the cultivar.
[15]
There are two other classification categories for cultigens, the
grex
[16]
and the
group
.
[17]
The
Code
then defines a
cultivar
as a "taxonomic unit within the classification category of cultivar". This is the sense of
cultivar
that is most generally understood and which is used as a general definition.
A cultivar is an assemblage of plants that (a) has been selected for a particular character or combination of characters, (b) is distinct, uniform and stable in those characters, and (c) when propagated by appropriate means, retains those characters.
[18]
Different kinds
[
edit
]
Which plants are chosen to be named as cultivars is simply a matter of convenience as the category was created to serve the practical needs of
horticulture
,
agriculture
, and
forestry
.
[19]
Members of a particular cultivar are not necessarily genetically identical. The
Cultivated Plant Code
emphasizes that different cultivated plants may be accepted as different cultivars, even if they have the same genome, while cultivated plants with different genomes may be regarded as the same cultivar. The production of cultivars generally entails considerable human involvement although in a few cases it may be as little as simply selecting variation from plants growing in the wild (whether by collecting growing tissue to propagate from or by gathering seed).
[20]
Cultivars generally occur as
ornamentals
and food crops:
Malus
'
Granny Smith
' and
Malus
'
Red Delicious
' are cultivars of apples propagated by cuttings or
grafting
,
Lactuca
'Red Sails' and
Lactuca
'Great Lakes' are lettuce cultivars propagated by seeds. Named cultivars of
Hosta
and
Hemerocallis
plants are cultivars produced by
micropropagation
or division.
Clones
[
edit
]
Cultivars that are produced asexually are genetically identical and known as
clones
; this includes plants propagated by
division
,
layering
,
cuttings
,
grafts
, and
budding
. The propagating material may be taken from a particular part of the plant, such as a lateral branch, or from a particular phase of the life cycle, such as a juvenile leaf, or from aberrant growth as occurs with
witch's broom
. Plants whose distinctive characters are derived from the presence of an intracellular organism may also form a cultivar provided the characters are reproduced reliably from generation to generation. Plants of the same
chimera
(which have mutant tissues close to normal tissue) or graft-chimeras (which have vegetative tissue from different kinds of plants and which originate by grafting) may also constitute a cultivar.
[21]
Seed-produced
[
edit
]
Some cultivars "come true from seed", retaining their distinguishing characteristics when grown from seed. Such plants are termed a "variety", "selection", or "strain" but these are ambiguous and confusing words that are best avoided. In general, asexually propagated cultivars grown from seeds produce highly variable seedling plants, and should not be labelled with, or sold under, the parent cultivar's name.
[22]
Seed-raised cultivars may be produced by uncontrolled pollination when characteristics that are distinct, uniform and stable are passed from parents to progeny. Some are produced as "lines" that are produced by repeated self-fertilization or inbreeding or "multilines" that are made up of several closely related lines. Sometimes they are
F1 hybrids
which are the result of a deliberate repeatable single cross between two pure lines. A few F2 hybrid seed cultivars also exist, such as
Achillea
'Summer Berries'.
Some cultivars are
agamospermous
plants, which retain their genetic composition and characteristics under reproduction.
[23]
Occasionally cultivars are raised from seed of a specially selected provenance ? for example the seed may be taken from plants that are resistant to a particular disease.
[24]
Genetically modified
[
edit
]
Genetically modified plants with characteristics resulting from the deliberate implantation of genetic material from a different
germplasm
may form a cultivar. However, the
International Code of Nomenclature for Cultivated Plants
notes, "In practice such an assemblage is often marketed from one or more lines or multilines that have been genetically modified. These lines or multilines often remain in a constant state of development which makes the naming of such an assemblage as a cultivar a futile exercise."
[25]
However, retired transgenic varieties such as the
fish tomato
, which are no longer being developed, do not run into this obstacle and can be given a cultivar name.
Cultivars may be selected because of a change in the
ploidy
level of a plant which may produce more desirable characteristics.
Cultivar names
[
edit
]
Every unique cultivar has a unique name within its denomination class (which is almost always the genus). Names of cultivars are regulated by the
International Code of Nomenclature for Cultivated Plants
, and may be registered with an
International Cultivar Registration Authority
(ICRA). There are sometimes separate registration authorities for different plant types such as roses and camellias. In addition, cultivars may be associated with commercial marketing names referred to in the
Cultivated Plant Code
as "trade designations" (see below).
Presenting in text
[
edit
]
A
cultivar name
consists of a
botanical name
(of a genus,
species
, infraspecific
taxon
, interspecific
hybrid
or intergeneric hybrid) followed by a cultivar
epithet
. The cultivar epithet is enclosed by single quotes;
[26]
it should not be italicized if the botanical name is italicized;
[27]
and each of the words within the epithet is
capitalized
(with some permitted exceptions such as conjunctions).
[28]
It is permissible to place a cultivar epithet after a common name provided the common name is botanically unambiguous. Cultivar epithets published before 1 January 1959 were often given a
Latin
form and can be readily confused with the
specific epithets
in botanical names; after that date, newly coined cultivar epithets must be in a modern vernacular language to distinguish them from botanical epithets.
[29]
For example, the full cultivar name of the
King Edward potato
is
Solanum tuberosum
'King Edward'. 'King Edward' is the cultivar epithet, which, according to the Rules of the
Cultivated Plant Code
, is bounded by single quotation marks.
[26]
For patented or trademarked plant product lines developed from a given cultivar, the commercial product name is typically indicated by the symbols "TM" or "®", or is presented in capital letters with no quotation marks, following the cultivar name,
[30]
as in the following example, where "Bloomerang" is the commercial name and 'Penda' is the cultivar epithet:
Syringa
'Penda' BLOOMERANG.
[31]
- Examples of correct text presentation:
- Cryptomeria japonica
'Elegans'
- Chamaecyparis lawsoniana
'Aureomarginata' (pre-1959 name,
Latin
in form)
- Chamaecyparis lawsoniana
'Golden Wonder' (post-1959 name,
English language
)
- Pinus densiflora
'Akebono' (post-1959 name,
Japanese language
)
- Apple 'Sundown'
- Some
incorrect
text presentation examples:
- Cryptomeria japonica
"Elegans" (double quotes are unacceptable)
- Berberis thunbergii
cv. 'Crimson Pygmy' (this once-common usage is now unacceptable, as it is no longer correct to use "cv." in this context;
Berberis thunbergii
'Crimson Pygmy' is correct)
- Rosa
cv. 'Peace' (this is now incorrect for two reasons: firstly, the use of "cv."; secondly, "Peace" is a trade designation or "selling name" for the cultivar
R.
'Madame A. Meilland' and should therefore be printed in a different typeface from the rest of the name, without quote marks, for example:
Rosa
Peace
)
Although "cv." has not been permitted by the
International Code of Nomenclature for Cultivated Plants
since the 1995 edition,
[32]
[33]
it is still widely used
[34]
and recommended by other authorities.
[35]
[36]
Group names
[
edit
]
Where several very similar cultivars exist they can be associated into a
Group
(formerly
Cultivar-group
). As Group names are used with cultivar names it is necessary to understand their way of presentation. Group names are presented in normal type and the first letter of each word capitalised as for cultivars, but they are not placed in single quotes. When used in a name, the first letter of the word "Group" is itself capitalized.
[37]
Presenting in text
[
edit
]
- Brassica oleracea
Capitata Group (the group of cultivars including all typical
cabbages
)
- Brassica oleracea
Botrytis Group (the group of cultivars including all typical
cauliflowers
)
- Hydrangea macrophylla
Groupe Hortensia (in French) =
Hydrangea macrophylla
Hortensia Group (in English)
- Where cited with a cultivar name the group should be enclosed in parentheses, as follows:
- Hydrangea macrophylla
(Hortensia Group) 'Ayesha'
[38]
Legal protection of cultivars and their names
[
edit
]
Since the 1990s there has been an increasing use of legal protection for newly produced cultivars.
Plant breeders
expect legal protection for the cultivars they produce. According to proponents of such protections, if other growers can immediately propagate and sell these cultivars as soon as they come on the market, the breeder's benefit is largely lost.
[39]
Legal protection for cultivars is obtained through the use of Plant breeders' rights and plant Patents but the specific legislation and procedures needed to take advantage of this protection vary from country to country.
[40]
Controversial use of legal protection for cultivars
[
edit
]
The use of legal protection for cultivars can be controversial, particularly for food crops that are staples in developing countries,
[41]
or for plants selected from the wild and propagated for sale without any additional breeding work; some people consider this practice
unethical
.
[42]
Trade designations and selling names
[
edit
]
The formal scientific name of a cultivar, like
Solanum tuberosum
'King Edward', is a way of uniquely designating a particular kind of plant. This scientific name is in the public domain and cannot be legally protected. Plant retailers wish to maximize their share of the market and one way of doing this is to replace the Latin scientific names on plant labels in retail outlets with appealing marketing names that are easy to use, pronounce, and remember. Marketing names lie outside the scope of the
Cultivated Plant Code
which refers to them as "trade designations". If a retailer or wholesaler has the sole legal rights to a marketing name then that may offer a sales advantage. Plants protected by
plant breeders' rights
(PBR) may have a "true" cultivar name ? the recognized scientific name in the public domain ? and a "commercial synonym" ? an additional marketing name that is legally protected. An example would be
Rosa
Fascination
= 'Poulmax', in which
Rosa
is the genus,
Fascination
is the trade designation, and 'Poulmax' is scientific cultivar name.
[
citation needed
]
Because a name that is attractive in one language may have less appeal in another country, a plant may be given different
selling names
from country to country. Quoting the original cultivar name allows the correct identification of cultivars around the world.
[43]
The main body coordinating plant breeders' rights is the
International Union for the Protection of New Varieties of Plants
(
Union internationale pour la protection des obtentions vegetales
, UPOV) and this organization maintains a database of new cultivars protected by PBR in all countries.
[44]
International Cultivar Registration Authorities
[
edit
]
An
International Cultivar Registration Authority
(ICRA) is a voluntary, non-statutory organization appointed by the
Commission for Nomenclature and Cultivar Registration
of the International Society of Horticultural Science. ICRAs are generally formed by societies and institutions specializing in particular plant genera such as
Dahlia
or
Rhododendron
and are currently located in Europe, North America, China, India, Singapore, Australia, New Zealand, South Africa and Puerto Rico.
[45]
Each ICRA produces an annual report and its reappointment is considered every four years. The main task is to maintain a register of the names within the group of interest and where possible this is published and placed in the public domain. One major aim is to prevent the duplication of cultivar and Group epithets within a genus, as well as ensuring that names are in accord with the latest edition of the
Cultivated Plant Code
. In this way, over the last 50 years or so, ICRAs have contributed to the stability of cultivated plant nomenclature. In recent times
[
when?
]
many ICRAs have also recorded trade designations and trademarks used in labelling plant material, to avoid confusion with established names.
[46]
New names and other relevant data are collected by and submitted to the ICRA and in most cases there is no cost. The ICRA then checks each new epithet to ensure that it has not been used before and that it conforms with the
Cultivated Plant Code
. Each ICRA also ensures that new names are formally established (i.e. published in hard copy, with a description in a dated publication). They record details about the plant, such as parentage, the names of those concerned with its development and introduction, and a basic description highlighting its distinctive characters. ICRAs are not responsible for assessing the distinctiveness of the plant in question.
[46]
Most ICRAs can be contacted electronically and many maintain web sites for an up-to-date listing.
[47]
See also
[
edit
]
Notes
[
edit
]
- ^
Cultivar
(
) has two meanings, as explained in
Formal definition
: it is a classification category and a taxonomic unit within the category. When referring to a
taxon
, the word does not apply to an individual plant but to all plants that share the unique characteristics that define the cultivar.
- ^
This ignored its prior use as a transitive verb in
Spanish
denoting "to farm, to cultivate, to grow, or to practice" (
Online Spanish dictionary
Archived
2010-01-11 at the
Wayback Machine
), and in
Portuguese
denoting
to cultivate, to husband, to farm, to plant, to polish, to reclaim, to improve
(
Ectaco online Portuguese dictionary
Archived
2023-04-07 at the
Wayback Machine
).
References
[
edit
]
- ^
Stanley J. Kays (3 October 2011).
Cultivated vegetables of the world: a multilingual onomasticon
. Springer. pp. 15?.
ISBN
978-90-8686-720-2
.
- ^
a
b
Bailey 1923
, p. 113
- ^
Spencer & Cross 2007
, p. 938
- ^
Lawrence 1953
, pp. 19?20
- ^
"HORT217 - Woody Landscape Plants"
.
Department of Horticulture and Landscape Architecture
- Purdue University
. Archived from
the original
on 2012-01-13
. Retrieved
2007-07-28
.
Also
Archived
2 August 2012 at
archive.today
- ^
Morton 1981
, pp. 38?39
- ^
Lawrence 1955
, p. 177
- ^
a
b
Lawrence 1955
, p. 180
- ^
Lawrence 1955
, p. 181
- ^
Lawrence 1955
, pp. 179?180
- ^
Trehane 2004
, p. 17
- ^
Lawrence 1960
, p. 1
- ^
Cultivated Plant Code Art. 2.3
Brickell 2009
, p. 1
- ^
Spencer, Cross & Lumley 2007
, p. 47
- ^
Cultivated Plant Code. Art. 2.1
Brickell 2009
, p. 6
- ^
Cultivated Plant Code. Art. 4
Brickell 2009
, p. 12
- ^
Cultivated Plant Code. Art. 3
Brickell 2009
, pp. 10?12
- ^
Cultivated Plant Code. Art. 2.2
Brickell 2009
, p. 6
- ^
Cultivated Plant Code. Preamble & Principles
Brickell 2009
, p. 19
- ^
Cultivated Plant Code, Article 2.20
Brickell 2009
, p. 9
- ^
Cultivated Plant Code, Articles 2.5?2.11
Brickell 2009
, pp. 6?7
- ^
Courses / RHS Gardening
Archived
2005-12-26 at the
Wayback Machine
- ^
Cultivated Plant Code, Articles 2.17?2.18
Brickell 2009
, pp. 7?8
- ^
Cultivated Plant Code, Articles 2.12?2.16
Brickell 2009
, pp. 7?8
- ^
Cultivated Plant Code, Articles 2.19
Brickell 2009
, pp. 8?9
- ^
a
b
Cultivated Plant Code Article 14.1
Brickell 2009
, p. 19
- ^
Cultivated Plant Code Recommendation 8A.1
Brickell 2009
, p. 15
- ^
Cultivated Plant Code Article 21.3
Brickell 2009
, p. 25
- ^
Cultivated Plant Code Art. 14
Brickell 2009
, p. 19
- ^
Dirr, Michael; Warren, Keith (2019).
The Tree Book: Superior Selections for Landscapes, Streetscapes, and Gardens
. Timber Press, Inc.
ISBN
978-1-60469-714-8
.
- ^
"
Syringa
'Penda' BLOOMERANG"
. Missouri Botanical Garden Plant Finder.
Archived
from the original on 2021-11-30
. Retrieved
2022-04-29
.
- ^
"Naming a Cultivar ?"
.
Australian National Botanic Gardens
- Botanical Web Portal
. 2000-07-28.
Archived
from the original on 2021-03-12
. Retrieved
2020-12-26
.
- ^
"How to name a new cultivar"
(PDF)
.
Food and Agriculture Organization
. 2020-06-10. Archived from
the original
(PDF)
on 2020-12-26
. Retrieved
2020-12-26
.
- ^
Chen, Guang; Liu, Chaolei; Gao, Zhenyu; Zhang, Yu; Zhang, Anpeng; Zhu, Li; Hu, Jiang; Ren, Deyong; Yu, Ling; Xu, Guohua; Qian, Qian (2018-01-05).
"Variation in the Abundance of OsHAK1 Transcript Underlies the Differential Salinity Tolerance of an
indica
and a
japonica
Rice Cultivar"
.
Frontiers in Plant Science
.
8
.
Frontiers Media
SA: 2216.
doi
:
10.3389/fpls.2017.02216
.
ISSN
1664-462X
.
PMC
5760540
.
PMID
29354152
.
- ^
"Horticulture 202 Laboratory 3"
.
Texas A&M University
General Horticulture
.
Archived
from the original on 2020-07-17
. Retrieved
2020-12-26
.
- ^
Finkle, Anita (2018-04-02).
"What is the proper way to write a botanical name (Latin name)?"
.
New York Botanical Garden
.
Archived
from the original on 2020-11-26
. Retrieved
2020-12-26
.
- ^
Cultivated Plant Code Art. 3
Brickell 2009
, pp. 10?12
- ^
Cultivated Plant Code Art. 15
Brickell 2009
, p. 19
- ^
P. Gepts (2004)
Who Owns Biodiversity, and How Should the Owners Be Compensated?
Plant Physiology 134, pp. 1295?1307
- ^
BSPB
Plant breeding ? The business and science of crop improvement
Archived
2011-09-11 at the
Wayback Machine
British Society of Plant Breeders booklet
- ^
"Adi, A.B.C.,
Intellectual Property Rights in Biotechnology and the Fate of Poor Farmers' Agriculture.
Social Science Research Network"
. 13 January 2005.
Archived
from the original on 2021-07-15
. Retrieved
2020-07-23
.
- ^
'Who owns nature?'
Archived
2011-09-11 at the
Wayback Machine
(article by nurseryman and plant hunter Michael Wickenden, published in
The Plantsman
)
- ^
Spencer, Cross & Lumley 2007
, pp. 76?81
- ^
Spencer, Cross & Lumley 2007
, p. 78
- ^
Cultivated Plant Code
Brickell 2009
, pp. 62, 67?83
- ^
a
b
See
International Cultivar Registration Authorities
Archived
2017-07-15 at the
Wayback Machine
- ^
Staff (2010).
"ISHS :: Commission Nomenclature and Cultivar Registration - International Cultivar Registration Authorities (ICRAs)"
.
ishs.org
. Archived from
the original
on 26 November 2014
. Retrieved
5 March
2011
.
Bibliography
[
edit
]
- Bailey, Liberty Hyde
(1923).
"Various cultigens, and transfers in nomenclature"
.
Gentes Herbarum
. 1 (Part 3): 113?136.
- Brickell, Chris D. (2009).
"International Code of Nomenclature for Cultivated Plants (ICNCP or Cultivated Plant Code) incorporating the Rules and Recommendations for naming plants in cultivation. 8th ed., adopted by the International Union of Biological Sciences International Commission for the Nomenclature of Cultivated Plants"
(PDF)
.
Scripta Horticulturae
.
10
. International Society of Horticultural Science: 1?184.
ISBN
978-90-6605-662-6
. Archived from
the original
(PDF)
on 2011-08-13
. Retrieved
2011-02-07
.
- Lawrence, George H.M. (1953). "Cultivar, Distinguished from Variety".
Baileya
.
1
: 19?20.
- Lawrence, George H.M. (1955). "The Term and Category of Cultivar".
Baileya
.
3
: 177?181.
- Lawrence, George H.M. (1957). "The Designation of Cultivar-names".
Baileya
.
5
: 162?165.
- Lawrence, George H.M. (1960). "Notes on Cultivar Names".
Baileya
.
8
: 1?4.
- Morton, Alan G. (1981).
History of Botanical Science: An Account of the Development of Botany from Ancient Times to the Present Day
. London: Academic Press.
ISBN
0-12-508382-3
.
- Spencer, Roger; Cross, Robert; Lumley, Peter (2007).
Plant names: a guide to botanical nomenclature. (3rd ed.)
. Collingwood, Australia: CSIRO Publishing (also Earthscan, UK.).
ISBN
978-0-643-09440-6
.
- Spencer, Roger D.; Cross, Robert G. (2007). "The International Code of Botanical Nomenclature (ICBN), the International Code of Nomenclature for Cultivated Plants (ICNCP), and the cultigen".
Taxon
.
56
(3): 938?940.
doi
:
10.2307/25065875
.
JSTOR
25065875
.
- Trehane, Piers (2004). "50 years of the
International Code of Nomenclature for Cultivated Plants
".
Acta Horticulturae
.
634
: 17?27.
doi
:
10.17660/ActaHortic.2004.634.1
.
External links
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edit
]
Look up
cultivar
in Wiktionary, the free dictionary.