Genus of tapejarid pterosaur from the Late Cretaceous
Bakonydraco
is a
genus
of
pterodactyloid
pterosaur
from the
Late Cretaceous
period (
Santonian
stage) of what is now the
Csehbanya Formation
of the
Bakony Mountains
,
Iharkut
,
Veszprem
, western
Hungary
.
Etymology
[
edit
]
Bakonydraco
was named in 2005 by paleontologists
Attila ?si
,
David Weishampel
, and
Jianu Coralia
. The
type species
is
Bakonydraco galaczi
. The genus name refers to the Bakony Mountains and to
Latin
draco
, "
dragon
". The
specific epithet
galaczi
honors Professor Andras Galacz, who helped the authors in the Iharkut Research Program, where fossils are since 2000 found in
open-pit mining
of
bauxite
, among them the remains of pterosaurs, the first ever discovered in Hungary.
Description
[
edit
]
Bakonydraco
is based on
holotype
MTM Gyn/3, a nearly complete
mandibula
, a fusion of the lower
jaws
. Also assigned to it, as
paratype
, is MTM Gyn/4, 21: parts from another jaw's
symphysis
(the front parts, having fused into a single blade-like structure, of the two lower jaws); azhdarchid
wing
bones and
neck
vertebrae
from the same area may also belong to it.
[1]
The lower jaws are toothless and the two halves of the mandibula are frontally fused for about half of its overall length, forming a long, pointed section that is compressed side-to-side and also expanded vertically, giving it a somewhat
spearhead
- or
arrowhead
-like shape from the side. This expansion occurs both on the lower edge and on the top surface, where the most extreme point corresponds with a transverse ridge which separates the straight back half of the symphysis from the pointed end in the front. The jaws of MTM Gyn/3 are 29 centimeters (11.4 inches) long, and the
wingspan
of the genus is estimated to be 3.5 to 4 meters (11.5 to 13.1 feet), which is medium-sized for a pterosaur. Because the jaws are relatively taller than other azhdarchids, and reminiscent of
Tapejara
, it could have been a
frugivore
.
[1]
Classification
[
edit
]
Initially,
Bakonydraco
was assigned to the family
Azhdarchidae
,
[1]
however, paleontologists Brian Andres and Timothy Myers in 2013 had proposed that
Bakonydraco
actually belonged to the family
Tapejaridae
, in a position slightly more
basal
than both
Tapejara
and
Tupandactylus
.
[2]
Indeed, the original paper describing this species compared the holotype jaw to
Tapejara
and
Sinopterus
,
[1]
implicating its affinities to this clade (or at least a large amount of
convergence
). If
Bakonydraco
is a tapejarid, it represents the only
Late Cretaceous
record of Tapejaridae known to date (aside from the slightly older
Caiuajara dobruskii
). A more recent phylogenetic study reinforces this placement.
[3]
The cladogram on the left follows the 2014
phylogenetic analysis
by Brian Andres and colleagues that also recovered
Bakonydraco
within the family Tapejaridae, more specifically within the tribe
Tapejarini
.
[4]
In 2020, in a phylogenetic analysis conducted by David Martill and colleagues,
Bakonydraco
was once again found within the Tapejaridae, this time consisting of two lineages: the
Tapejarinae
and the
Sinopterinae
,
Bakonydraco
was recovered within the subfamily Sinopterinae in the basalmost position, unlike in the analysis by Andres and colleagues. Their cladogram is shown on the right.
[5]
Topology 1
: Andres
et al.
(2014).
|
Topology 2
: Martill
et al.
(2020).
|
See also
[
edit
]
References
[
edit
]
- ^
a
b
c
d
Osi, Attila; Weishampel, David B.; Jianu, Coralia M. (2005).
"First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary"
(PDF)
.
Acta Palaeontologica Polonica
.
50
(4): 777?787
. Retrieved
July 28,
2009
.
- ^
Andres, B.; Myers, T. S. (2013). "Lone Star Pterosaurs".
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
.
103
(3?4): 383?398.
doi
:
10.1017/S1755691013000303
.
S2CID
84617119
.
- ^
Wu, W.-H.; Zhou, C.-F.; Andres, B. (2017).
"The toothless pterosaur
Jidapterus edentus
(Pterodactyloidea: Azhdarchoidea) from the Early Cretaceous Jehol Biota and its paleoecological implications"
.
PLOS ONE
.
12
(9): e0185486.
Bibcode
:
2017PLoSO..1285486W
.
doi
:
10.1371/journal.pone.0185486
.
PMC
5614613
.
PMID
28950013
.
- ^
Andres, B.; Clark, J.; Xu, X. (2014).
"The earliest pterodactyloid and the origin of the group"
.
Current Biology
.
24
(9): 1011?6.
doi
:
10.1016/j.cub.2014.03.030
.
PMID
24768054
.
- ^
Martill, David M.; Green, Mick; Smith, Roy; Jacobs, Megan; Winch, John (April 2020). "First tapejarid pterosaur from the Wessex Formation (Wealden Group: Lower Cretaceous, Barremian) of the United Kingdom".
Cretaceous Research
.
113
: 104487.
doi
:
10.1016/j.cretres.2020.104487
.