?? ????????? ??????? ??? ???? ??????? ????????? ??????? ?????? ????? ?????? ????????? ????? ????
???? ????????
?????
???
??????????
? ????
????????
??
????
(brain) ?????? ????
????????? ??????
??? ?????????
?????????? ??????
?
??????? ??????
? ??????????? ??? ???
???
??? ??????? ???? ????????? ?????????? ?????? ????
???'???
,
???????
,
??????'?
???
????????
? ???????? ??????; ??? ???? ???
??????? ??????
(nerve net) ????? ???????? ???? ???? ???????? ???????? ?????? ??? ???? ????, ????, ???? ?????, ????? ????, ????? ???? ??? ?????? ?????????? ???? ????? ?????????? ??????? ???? ????????? ??????? ???? ???? ??? ?????? ?????? ????? ????????? ????????? (??????? ???? ???) ?????? ??-?? ?????
??????????
?????
[1]
?? ????????
????????????????
?? (synapse) ?????-????? ??? ??? ?? ????? ????? ????????? ??? ??????? ?? ????? ?? ?????????????? ????? ???? ???? ????'????????? ??????????? ????? ????-?????? ???? ?????? ????????
??? ????
(action potential) ?????? ????????? ?? ??? ?? ????? ?????? ????? ???????? ?? ?????
??????????
?????? ????? ??? ???? ???? ?? ???????? ?????????????? ?????????? ???? ?????? ????? ??????????? ????? ??
???'?
???? ????????? ??????? ????????? ???? ?? ???? ??? ???? ??????? ???? ?'?? ?? ?????????? ??????????? ???? ??????? ??? ??????? ?????? ??????? ???? ???????????? ?? ??????????
??????????????
???
?????? ???????
? ??? ??????? ???????? ??????? ????? ???, ?????? ???? ???????? ???? ?????? ??? ????? ????????-???????? ??????????? ???? ????? ??????? ???????? ???? ??? ????? ?????????
????????? ??????? ?????????? ???????????? ?????? ???? ?????? ?????? ??????? ???? ?????? ????? ???? ??????? ???? ?????????? ??????? ?????? ???? ????
[2]
?????? ??????????????? ???????? ?????????? ????? ?????
?????????
???? ???? ??? ??? ???????????? ??????????? ???????????? ??? ????, ?????? ????????? CPU-? ??? ??????? ????????? ??? ???? ???? ??? ??? ??? ???? ???? ???????? ????
??????? ?'?????????? ?????? ????? ????????? ???? ??? ????? stain ??? ????: ??? ??? ??????????? ??? ???? ???
????? ??????
????? ?????? ?????? ??????????
????? ????? ???????? ????????? ??? ??? ????? ?????? ???????? ??? ??? ??? ????????? ?????? ?????? ????
[3]
?? ????, ????????? ????????? ????????? ???? ??????? ???? ?????? ????
[4]
????????? ???? ??????? ??? ???? ???????? ???????? ???;
[5]
?????? ??????? ???????? ???? ????? ??? "????" ???????? ????? ???? ???, ?? ?????????? ??? ?????????? ??????? ???? ????
[3]
???? ????? ????? ???? ???? ??? ??????? ??? ???? ????- ??? ??? ???????
???? ???
(grey matter) ??? ???? ????
????? ???
(white matter)? ????? ????? ????? ????????? ??? (stain) ??????? ??? ????????? ???? ???? ???????? ??? ????? ?????
????????? ??????
??? ????? ???? ??? ??????? ???? ???? ??? ????? ??? ???? ??? ?? ???? ????? ??? ????? ?????
[6]
??????????? ?????? ??? ????????? ????? ?????? ?????? ???????? ???? ?? ??????? ????
????????????????
????? ??? ??? ???? ??????? ?????? ?????? ??? ?? ?? ???? ?????? ?????? ?? ???? ??????? ????????? ????????????? ?????? ???????
???? ????????? ????????? ??? ??????? ?????? ??? ????:
??????????
???
????'???????
(neuroglia)? ????'??????? ?? ??????? ????????????? ??? ?????? ??????, ?????, ????? ??? ??????? ????? ????? ???? ?????? ??????????? ????????? ??????? ???????????? ??? ???? ???? ??? ????
[7]
??? ???? ??? ?????? ????? ????? ??????? ??? ??????? ???? ?????????? ??????
[7]
?????? ????? ???? ????'????????? ??????????? ????? ?????? ???? ?????? ??????? ??? ???? ?'? ?????
[8]
????????? ????????
??? ????
(action potential) ???? ???????-????????? ?????? ????????? ??? ??????? ?? ???????? ?????? ???? ???????? ?? ???? ???? ??? ?????? ?????? ?-??? ?????/??????? ????? ?????
[9]
?????? ?? ????????????
????????????????
(synapse) ???? ????? ???????? ????? ?????? ????
[7]
?????? ?????? ??? ??? ???? ??? ???????????????? ??? ???? ??? ???
??????? ??????
(neurotransmitter) ???? ????????? ?????? ?????? ???? ?? ??????? ?????? ?? ?????? ???? ?????? ?????? ?? ?????
[7]
Neurons often have extensive networks of
dendrites
, which receive synaptic connections. Shown is a
pyramidal neuron
from the hippocampus, stained for
green fluorescent protein
.
???????????????? ????? ???????????? ???????? ???????
[10]
????? ??????????? ????? ???? ??????? ???? ??????? ?????? ??? ????????????????? ???????? ?? ??????? ???? ?????? ????? ?????? ??? ????????? ???????????????? ??? ???? ???? ??? ????;
[11]
???? ???? (fruit fly) ????? ????? ????? ????? ???????????????? ?????
[12]
????????????????? ??? ??? ???: ???????? ???? ??????? (excitatory) ???? ??? ???; ??????? ?????? (inhibitory); ????????
???????? ?????? ???????
(second messenger system) ??????? ??? ???? ?? ?????? ???? ?????????? ????????? ?????????? ????? ??????? ?????? ??????? ??? ??? ?? ????????????????? ?????-????????? ???????? (synaptic plasticity) ?????? ??? ??????? (learning and memory) ???????? ???????????
[10]
????? ??????? ??? ?????? ?????? ??? ??? ??????? ???? ??? ??? ???????????? ??? (nerve fiber tracts) ?????? ???? ??????? ???????
???????? ????
???? ????? ???? ???? ?? ????? ??????? ??? ??????? ????? ???? (????????????? ?????? ????)? ???????? ??? ???? ???? ???? ???????????? ??? ????? ???? ???? ???????
????? ???
??? ???, ??? ??????????? ??? ??? ??? ????? ???? ??? ???????
???? ???
??? ????
[7]
- ↑
Pelvig, DP; Pakkenberg, H; Stark, AK; Pakkenberg, B (2008). "Neocortical glial cell numbers in human brains".
Neurobiology of Aging
????
29
(11): 1754?1762.
doi
:
10.1016/j.neurobiolaging.2007.04.013
.
PMID
17544173
.
- ↑
Yuste, Rafael; Church, George M. (March 2014).
"The new century of the brain"
.
Scientific American
????
310
(3): 38?45.
doi
:
10.1038/scientificamerican0314-38
.
PMID
24660326
. Archived from
the original
on 2014-07-14
.
https://web.archive.org/web/20140714190157/http://www.columbia.edu/cu/biology/faculty/yuste/Publications/Yuste_Church_SciAm14.pdf
? ???? ??? ????: 2015-07-01
.
- ↑
3.0
3.1
Shepherd, GM (1994).
Neurobiology
. Oxford University Press. ??????. 3.
ISBN
978-0-19-508843-4
.
- ↑
Sporns, O (2010).
Networks of the Brain
. MIT Press. ??????. 143.
ISBN
978-0-262-01469-4
.
https://books.google.com/books?id=v1DBKE7-UrYC&pg=PA143
.
- ↑
Ba?ar, E (2010).
Brain-Body-Mind in the Nebulous Cartesian System: A Holistic Approach by Oscillations
. Springer. ??????. 225.
ISBN
978-1-4419-6134-1
.
https://books.google.com/books?id=NAbMHo-ux58C&pg=PA225
.
- ↑
Singh, I (2006).
"A Brief Review of the Techniques Used in the Study of Neuroanatomy"
.
Textbook of Human Neuroanatomy
. Jaypee Brothers. ??????. 24.
ISBN
9788180618086
.
https://books.google.com/books?id=bBMS013dmycC&pg=PA24
.
[
?????????? ???
]
- ↑
7.0
7.1
7.2
7.3
7.4
Kandel, Eric R.; Schwartz, James Harris; Jessell, Thomas M. (2000).
Principles of neural science
. ??????? New York: McGraw-Hill.
ISBN
978-0-8385-7701-1
.
OCLC
42073108
.
- ↑
Douglas, RJ; Martin, KA (2004). "Neuronal circuits of the neocortex".
Annual Review of Neuroscience
????
27
: 419?451.
doi
:
10.1146/annurev.neuro.27.070203.144152
.
PMID
15217339
.
- ↑
Barnett, MW; Larkman, PM (2007). "The action potential".
Practical Neurology
????
7
(3): 192?197.
PMID
17515599
.
- ↑
10.0
10.1
Shepherd, GM (2004). "Ch. 1: Introduction to synaptic circuits".
The Synaptic Organization of the Brain
. Oxford University Press US.
ISBN
978-0-19-515956-1
.
- ↑
Williams, RW; Herrup, K (1988). "The control of neuron number".
Annual Review of Neuroscience
????
11
: 423?453.
doi
:
10.1146/annurev.ne.11.030188.002231
.
PMID
3284447
.
- ↑
Heisenberg, M (2003). "Mushroom body memoir: from maps to models".
Nature Reviews Neuroscience
????
4
(4): 266?275.
doi
:
10.1038/nrn1074
.
PMID
12671643
.
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