?? ?????????? ???????? ?????
?????
??? ?????? ???
(
???????????
:
Sir Paul Nurse
)? (???? ?? 25 ????? 1949)? ??
???? ?????
???????
??????? ?????? ??????? ???????
??????? ???????? ????? ???? ??????? ????.
[18]
[19]
[20]
???
?????? ???? ?? ??? ????? ??????? ?? ????
??? 2001 ???????? ??
?????? ???????
???? ????
??????? ????????? ?????? ???????? ???? ????? ?? ?????? ??????? ??
???? ??????
.
[21]
????? ????? ???????
[
???
]
????? ????? ??? ??
????
???
?????
?
?????
????? ?? ??????? ??????? ????? ??? ?????? ??? ??????. ???? ???????? ?????? ???? ????? ?????? ??????? ?????? ????? ????.
[22]
???? ??? ?? ??? ???? ?? ???? ??? ????.
[23]
???? ?????? ?? ?????
???????
?????? ?????? ?????? ???????. ???? ??? ???????
?????? ???????
?????????
??????
?????
???? ?? ????? ????? ???????? ???????? ?????? ??? ????? ?????? ??????? (?? ?? ??). ???? ???? ???? ??
????? ????????
???? ?? ???? ?????? ??
????? ????????
?? ???? ?????. ??? ??? ???? ??????????? ??
??? ???????
??? 1970 ??
????? ????????
[24]
?????
?????????
?? ??? 1973 ??
????? ???? ??????
????? ?? ????????? ????? ????????.
[25]
?? ???? ???? ?? ??? ????????? ??
????? ???
?????? ??????
?????? ?????
.
????? ??????? ???????
[
???
]
???? ???
???? ?? ??? ?????????
?? ????? ????? ???????? ??
????? ??????
??????? ???? ??????? (1973-1979).
[26]
[27]
??????? ?? ??? 1976? ??? ??? ????? ?? ?? ?? 2 ??
?????? ?????
.
[28]
[29]
????? ??? ????? ?? ????
???? ??????
??
??????? ?? 1
???
??????? ??
????????? ??
??????? ?? 2
??? ?????
???????? ????????
. ?? ??? 1987? ??? ??? ????? ???????? ?? ???????? ?? ?? ??? 1 ????? ???? ??? ?????? ??????? ??????? ??? ????????.
[30]
????? ????? ??????? ??? ?????? ????? ??? ????? ???? ?? 1 (?????)? ?? (????????)? ?? 2 (?????)? ??? (???????? ????????). ????? ??? ???????? ???? ???????? ?????
????????
???????? ???????? ??? ???????? (?? ?? ???)? ?????? ??????? ?? ???????? ?? ????? ??? ????. ???? ??? ?????????? ???? ??????? ????? ????? ?? ??? ???? ?????? ?? ?????? ???? ????. ??? ?? ????? ?????? ???? ????? ??????? ?????????? ?????? ???????? ???? ????? ???? ???? ??????. ??? ?????? ?????? ???? ??? ???? ????? ??? ??? ??????? ????? ?? ???? ??????? ?????? ????? ????.
[31]
?? ???? ????? ??? ?????? ?????? ??? ??? ????? ?? ?? ?? 2? ???? ????? ?? ???????? ?? ????? ?? 1 ??? ????? ??? ????? ???? ?????? ????????? ???? ????? ??????? ??? ????? ?? 2 ??? ????? ??? ????? ????? ??????. ??? ??? ?????? ?? ??????
?????? ??
???? ???? ?? ??? ?????????? ????? ??????? ????? ?? ?? ??? 1? ?? ?????. ????? ??? ??????? ????? ?????? ?????? ????? ???????? ??? ???????? (?? ?? ???) ?? ???? ????? ?? ????? ?????? ????????.
[31]
????? ??????
[
???
]
?????
[
???
]
- ^
Encyclopædia Britannica | Sir Paul M. Nurse
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- ^
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.
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,
QID
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??????? ??? ???
??????? ?????? ?????
?????
?????????????? ??? ???? ????? ??
???????
.
(???????? ?????)
- ^
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. Nobelprize.org.
???? ??????
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- ^
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, BBC
???? ??????
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"
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?????
?? 2020-04-14
. ???? ???? ??????
2017-12-10
.
- ^
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???? ??????
21 ????? 2015 ??
Wayback Machine
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.
????:EThOS
. ????? ??
?????
?? 2020-07-29.
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??? ???
??? ????
????? ????
? ?????? ?? 28 ????? 2020 including the Nobel Lecture Nobel Lecture December 9, 2001
Controlling the Cell Cycle
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.
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.
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.
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?
?
The Nobel Prize in Physiology or Medicine 2001
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???? ??????
2020-07-29 ??
Wayback Machine
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