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عصر كمبريعصر أردوفيسيعصر سيلوريعصر ديفونيعصر كربونيعصر برميعصر ثلاثيالعصر الجوراسيعصر طباشيريباليوجيننيوجين
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عصر كمبريعصر أردوفيسيعصر سيلوريعصر ديفونيعصر كربونيعصر برميعصر ثلاثيالعصر الجوراسيعصر طباشيريباليوجيننيوجين
???? ????? ??????? ?????? ???? ????? (??? ??? ????) ?????? ????????? ??????? ???????? ???? ?? ???? ????? ?????. ??? ???? ???? ??????? ???????? ?? ??? ???? ????? ??????. ???????? ?????? ???????? "?????? ??????" ?? ????? ????? ?????? ????? ?? ???????? ???? ????? ???????? . ( ??????? ?????? ??????? )

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96% ?? ??????? ???????
70% ?? ?? ????? ????????? ???????.

?????? ????? ?????? ??????? ??????? ???? ????? ?????? [3] ?? ?????? ????? ??? ??? 251.9 ????? ???? [4] ?? ?????? ??? ??????? ??????????? ?????? ???????? . ??? ??? ????? ???? ???? ?????? ?????? ?????? ????? ??? ??? ??? 96% ?? ?? ????? ??????? ??????? ? [5] ??????? ??? 70% ?? ?? ????? ????????? ???????. [6]

??? ???? ??? ???????? ?? ???????? ??????? ?????? ??????? ??????? . [7] ?????? ??? 57% ?? ?? ???????? ?83% ?? ???? ???????. ????? ???? ?????? ?? ?????? ?????????? ?????? ????? ????? ???? ????? ?????? ??? ????? ??????? ?????? ???????? ??????. [5] ??? ???? ??? ????? ???? ≪?? ?? ?????????? ???????≫. [8]

????? ???????? ?? ??? ???????? ?? ?? ???? ????? ??? ???? ?????. [4] [6] [9] [10] ???? ?????? ?? ??????? ???????? ????? ??? ????????? ???? ????? ???????? ???? ???? ???? ???? ??????? ?? ??? ??????? ??????? ???? ??? ???? ?????. ??????? ??????? ???? ?????? ???? ???? ?? ????? ??????? ?? ????? ???????? ?????? ?????? ????? ?? ????? ??????????? ?? ????? ??????? [11] ? ??????? ??? ??????? ?? ??? ???????? ?????? ????? ????? ??????? . ???? ????? ???????? ????????? ???? ????? ??? ????? ????? ??? ?? ???????? ?????? ??????? ??????? ?? ????? ???????? ???? ???????? ????????.

????? ???????? [ ??? ]

??????????? ??????? [ ??? ]

????? ??????????? ??????? ???? ??????? ???? ?????? ????? ?????? ???????. ???? ??? ???? ??? ??? ?? ????? ?? ????? ???? ????? ??? ???? ????? ?????? ???????. ??? ????? 286 ?? ??? 329 ????? ?? ??????????? ??????? ???? ????????? ???????? ????????? ?????? ?????? ??? ???????? ??????? ?? ????? ??????. ????? ??? ???????? ?? ?????? ?????? ?? ????? ???? ?? ????? ???????? ? ????? ?? ?????? ?? ????????. [12] [13]

??? ???????? ?? ?????? ????? ??? ???????? ????? ??? ??????? ??????? ??????? ?? ??????? ?????????? ????? ??? ???? ????? ??? ??????? ????? ?? ???? ????? ??????? ?????? ??????? ???????. ???? ??? ???????? ????? ???? ???????? ???? ???????? ??????? ?? ????? ???? ????? ??????? ?? ?????? ????? . [14]

??? ???????? ???????? ???? ??? ???????? ?????? ???????? ???????? ???????? ???? ?? ???? ?????? ?? ????? ??????? ??? ????????? ???? ????? ???? ?????? ????? ????? (??????? ??????? ??? ???? ????? ?????). ??? ???? ?????? ???????? ??????? ???????. [15]

????? ??????? ??????????? ??????? ??????? ??? ??? ?????? ?????? ?????? (??? ???? ??? ????)? ????? ???? ???????? ?????? ?? ??????? ??? ???????? ?????? ???????? ??? ???????????? ????????? ? (????? ?????)? ???? ???? ?? ????? ?????? ????? ????? ??????? ???? ???.

???? ????????? ???? ?????? ?????? ??? ??? ?????? ???? ????? ?????? ??????? ????? ?? ???????? ?????? ????? ??????? ???????? ??????? ??????? ???? ???????? ????? ?????? ?????? ??? ????? ?????? ?????? ?? ????? ???? ?? ???? ???????. ?? ???? ?????? ??????? ??? ?????? ???? ????????? ??????? ???? ??? ????? ?????? ?? ????? ????? ??????.

???? ????????? ???? ???? ?? ???? ????? ???? ????? ???? 30 ????? ??? ??? ?????? (????? ????? ??????)? ?? ?????? ??????? ??? 10 ?????? ??? ?? ????? ???????? ?? ????? ????? ?????????. ?? ??? ???????? ??????? ???? ??? ???? ???? ?? ???????? ?? ?? ?????? ???????? ??????? ????????? ???? ??????? ??????? ?? ???????? ??? ?? ????. ????? ?????? ???????? ???? ??? ???? ????? ??????- ???????. ??? ???????? ??(????? ??????-???????) ??? ???????? ??? ???? ?? ???? ???????. ???? ????? ???????? ????? ?????? ?????? ??? ??????? ?? ???????. [16]

???? ???? ??????????? ???? ????? ????????? ?? ???? ??????? ?? ??????? ?? ??????? ????????? ???? ??????? ?? ???? ????? ??????. ??? ???? ?????? ?? ?????? ?? ????? ???????? ???? ????? ???????? ?????  ?????? ?????? ??? ????? ???? ??? ????? ??????????? ???? ????? ??? ?????? ???????. [17]

??????????? ??????? [ ??? ]

??? ??? ????? ?????? ???? ???? ?? ??????? ?????? ??????????? ??????? ??? ?? ??? ???? ??????? ???????? ??? ???????. ???? ????? ????? ?????? ???? ?????? ????? ????? ???????? ?????? ???? ???????? ??? ???????? ??????? ?????? ???????. ?????? ?????? ?? ???? ??? ????? ?????? ???? ???? ???? ???? ????. ???? ?????? ??????? ??????? (??? ??????? ???? ????? ????? ????? ????? ?????) ?? ???????? ???? ????? ????? ??????? ??????? ??? ?????????? ?????? ?? ????????. ??? ???????? ?????? ?? ????? ????? ?????? ????? ?? ??? ???? ??????? ???????? ??????? ???????? ??????. [18]

???? ??????? ??????? ????????? ???? ???? ????? ??? ???? ????? ??????-??????? ????? ???????? ?????? ?? ??? ???? ???? ????? ?? ???: ??? ??? ??????? ????? ???? ?????? ???????? ?? ????? ?? ??????? ?? ??? ???????? ??? ???????????????? ?????????????? ????????? ??????? ????????. ?????? ?? ?? ??????? ???????? ???????? ?????? ??????? ??????? ???????? ?????????? ??????????? ?????? ?????? ????? ??????. ??? ????? ????? ????? ??????? ??????? ??????? ?????? ????? ???????? ???? ???? ?? ??????? ??????? ????????? ???????. [19]

???????? ??????? [ ??? ]

??????? ?????? ?????? ?????? [ ??? ]

????? ????????? ???????? ??????? ?????? ?????? ?? ?????? ??? ?????? ???? ?????? ?????????. ???????? ????? ?????? ?? ???????? ???????? ?? ????? ???? ?????????? ???????? ?????? ≪??? ???????≫ ??? ????? ???????. ???? ???????? ??????? ?? ???? ??????? (????? 50%) ?? ???? ?????? ?? ????? ??? ???????? ?????????. ??? ???? ???? ????? ????? ????? ????? ???????? ?? ?????? ????? ?? ???? ???????? ?????????? ??????? ???? ??????? ???????? ??????? ??? ?????? ???? ?????? ??????? ?? ??? ????????.

??? ???? ????? ??????-???????? ????? ????????? ??????? ????????? ?? ???? ?????? ?? ??????? ???????? ?????? ??? ????? ?????? ??? ??????? (?????? ??????)? ??????? ??????? (??????? ?????). ??????? ?????? ?????? ?????? ??????? ??? ?????? ?????? ????? ????? ????? ?????? ??????? ??????? ????????. [20]

???? ?????? ??? ????? ?? ?????? ???? ?????? ?? ??? ???????? ?????? ?????? ???????? ???? ????? ???? ???? ???????? ??? ???? ????? ????? ??????? ??????? ?????? ??? ???? ?????. ?? ??? ????? ???? ?????? ??? ????????? ??????? ??????? ??????????? ???? ??? ??????? ??????? ??????? ??????? ?????? ?? ???? ???????? ??????? ?????? ??? ?? ??? ???????? ???????? ??? ?? ????? ?????????. ?? ??? ????? ????? ??????? ???? ?? ?????? ?????? ?????? ??? ???? ?????? ???? ??? ???? ?? ??? ????. ???? ??? ???????? ?? ???????? ??????? ??? ???? ???? ???? ???????? ??????. ???? ??? ???????? ?? ???????? ??????? ??? ??????? ??????? ???????? ??? ???? ????? ????? ??? ??? ????? ???? ????? ?????? ??????? ??????? ???????. ?? ?????? ????? ??????? ????????? ?? ????????? ???????? ?? ?????? ?? ??? ???? ?? 13 (δ13C) ?????? ???? ??? ????? ?????. ??????? ??????? ????? ?????? ?????? ?? 4 ??? 5 ?????? ???. [21]

???? ????? [ ??? ]

?? ???? ????? ????? ?????? ?? ????? ????? ???????? ???? ???????? ?? ????? ??????? ?????? ????? ???????. ????? ≪???? ?????≫ ???? ???. ??? ???? ??? ?? ???????? ???????? ?????????? ??????? ?????? ??????? ?? ????? ????? ??????? ????? ?? ???????. ???? ??? ???????? ????? ?? ????? ????? ?? ??????? ????? ???????? ??? ????? ????? ??????? ????? ?????. ???? ?????? ?? ???? ???? ???????? ???? ???? ????? ?? ?????? ???? ?????? ????? ?????? ???????? ??? ????? ?????? 10 ?????? ??? ?????? ?????? ????? ?? ???????? ?? ?????? ?????? ???????? ????? ????. ??????? ??????????? (??????? ???? ?? ?????? ???????? ?????)? ??? ?????? ???? ??????? ?? ????? ?????? ??????? ????????? ?? ???? ?????? ?????. [22]

?? ????? ????? ?? ???? ??? ????? ???? ???? ??????? ???????? ?? ????? ????? ???????. ????? ???? ????? ??????? ??????? ?????? ????? ?? ????????? ?? ?????? ??? ??????? ???? ??? ????? ???? ??? ????? ?? ????? ????? ???????. ??? ??? ????????? ??? ???? ??????? ?? ?????? ?????? ?? ??????? ????? ?????? ?? ???? ?????? ?????? ???? ?? ??? ??????. ????? ?? ???? ?? 95% ?? ??? ???????? ??????? ???? ?????? ??? ???? ????? ?????? ???????? ?????? ????? ????? ?? ???????? ??????????? ???? ????? ??? ???? ????? ?????? ???????. [23]

???? ????? [ ??? ]

????? [ ??? ]

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